Marvel Database

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Skrulls under the command of Sl'gur't

The Skrull race is a species of green skinned humanoids that originate from the planet Skrullos in the Andromeda Galaxy many eons ago.

Their earliest origins and evolution is lost to time. Xorr the God-Jewel has claimed that the Skrulls, along with most other humanoid races are the descendants of the ancient Xorrians.[1] Another group called the Builders have also claimed to be responsible for creating the Skrulls and other races.[2] However these claims are unsubstantiated. Whatever the case, the Skrulls—like many other races in the universe—were selected by the enigmatic Celestials for experimentation, creating three distinct Skrull variants: baseline Skrulls with no inherent abilities but having the capacity for mutation, the neigh powerful Skrullian Eternals, and Skrullian Deviants.[3][4]

Yet another offshoot developed from the Skrulls, a race of black magic practitioners who became known as the Dire Wraiths, who soon departed Skrullos to inhabit the Dark Nebula and espoused plans for universal domination.[5]

The Deviant of Armechadon known as Tantalus, created by the rogue Celestian known as the Dreaming Celestial, convinced Deviants of all other races to eliminate their baseline and Eternal counterparts, including those of the Skrull race.[6] Whatever the case, millions of years ago,[citation needed] the Deviant Skrulls went to war against the other offshoots of their race, under the command of the warrior princess Sl'gur't until the vast majority of the other Skrull races were wiped out. At the time they believed to had drove the other races to extinction except for a lone Eternal Skrull named Kly'bn.

The ascension of Sl'gur't and Kly'bn to godhood

When Sl'gur't's forces stormed the "Unworthies"'s stronghold in the Valley of the Esul, Kly'bn convinced Sl'gur't not to kill him, as doing so would ultimately mean the death of their race and their greater destiny. Kly'bn spoke of a destiny beyond the stars where the Skrull people would rule over all those who were inferior Skrulls. Accepting this truth, Sl'gur't and Kly'bn were married, and transcended into godhood. This became the foundation of the Deviant Skrull's religion, they formed a religion called the Book of Worlds that was based on a manifest destiny over the entire universe. Sl'gur't and Kly'bn were worshiped as gods to the Skrulls, and over the centuries that followed the religion endured, however many of the Skrulls believed the tales from the Book to be myth, but devout followers endured over the ages. These devout worshipers formed a religion called the Dard'van.[7]

Nogor in the aftermath of an ancient Skrull battle

Over the centuries, the worlds invaded by the Skrulls the various pantheon gods of the 978 other worlds were either killed or enslaved. Among those gods, Hadith, Omen-Maker, Path-Ligher the god of the Queega of Quolan, Blessed-of-Litters of the Druff of Ryas, and Ceffyad the Righteous of Idoidea Swarm Collective had been co-opted and enslaved by the Skrull gods. Statues commemorating their fall and assimilation into Skrull culture soon became much of the imagery used in Skrull places of worship.[8] One of the most powerful and longest living Skrulls was Nogor, who was the talisman of the Skrull people and a step below the gods himself. Nogor often was present in the vicinity of Skrull conflicts as a show of approval for the Skrull gods.[9]

The opening strike of the Kree/Skrull War

Ancient History

About 10 million years ago, the Skrull became a great technologically advanced empire and a peaceful race for a time, they established colonies all over the Andromeda galaxy, establishing a new throneworld on Tarnax IV. Under the rule of Emperor Dorrek I, the Skrulls sought to share their technological advancement with another race. They discovered the planet Hala home to two distinct races: the blue skinned humanoids known as the Kree and the plant-like Cotati. Unwilling to share their gifts with two races, the Skrulls placed a number of each race on Blue Area of Earth's moon with the challenge to see who could build the most advanced city within a year. When the Skrulls returned they found that the Kree used Skrullian technology to build a massive technological metropolis, while the Cotati managed to breath new life on the surface of the moon, creating a lush forest. The Skrulls ruled in favour of the Cotati, angering the Kree who slaughtered Dorrek and his part. They then reversed engineered Skrull technology to build their own warships and attack the Skrull homeworld. This act of aggression led to the beginning of a the centuries long Kree/Skrull War, the Skrulls abandoned their peaceful ways and became a race of conquerors taking over planets by force and expanding their galactic empire.[10][11][12] Around this time, a Skrull ventured into the Citadel of Light and Shadow on a quest to obtain the reality altering device known as the Crystal of Ultimate Vision. The Skrull managed to get to the Crystal, and sought to claim it for his own. That quest was in fact a test, a mature species renouncing that power, and the Skrulls therefore failed the test: The Skrulls ended up in an evolutionary dead end and the Skrull who failed the test was turned into a statue that would go on to stand before the Citadel.[13]

Average Skrull male

Thousands of years ago, the Skrulls were involved in a scientific endeavor to invent the Lifestone Tree a collection of powerful gems that endowed the Chosen Eight of Fate to be guardians of the galaxy, one of their number included a Skrull.[14] The Chosen Eight were active for about 200 years. During this time four members died in battle, two of died of old age, one retired, and the final member's whereabouts became a mystery for centuries. The fate of the Skrull who fought with the Chosen eight is unknown. Ultimately, the gems making up the Lifestone Tree were lost after an attack by space pirates and the gems were lost on Earth well into more modern times.[15] Relatively around this time another Skrull was responsible for creating a Cosmic Cube a powerful device that could alter reality to the whims of the user. When given to the Skrull emperor at the time he used the power of the Cube to make himself a god. However the Cube gained it's power through the realm of the Beyonders and soon developed a megalomaniacal persona that matched its user and laid waste to the entire Skrull empire, destroying many of its world's and reducing the Skrulls to barbarism. Eventually the Cube matured changing itself into the so-called Shaper of Worlds and left to seek those with imagination enough to utilize his power, leaving the Skrulls to rebuild their crippled empire[16].

Average Skrull female.

The Skrulls rebuilt their empire over the centuries and continued age old war with the Kree and continued to expand their empire. They soon became aware of Earth, a planet that was strategically between the Kree and Skrull galaxies and this world became the focal point of many invasion plots by the Skrulls to gain it as a tactical advantage in their war[10]. Presumably, hostilities between the Kree and the Skrulls cooled around this time, and it is possible that it was at this time the Convention of Fornax was signed[17] which among other probably provisions included a moratorium on torturing prisoners of war.

Early Interactions with Earth

Skrull Cows circa 1776

The earliest known interaction between the Skrulls and the planet Earth dates back to the Spanish Inquisition at some time between the late 1400s or early 1500s. The religious natives who found the corpses believed the Skrulls to be demons and cast their bodies in an inaccessible ravine. While persecuting a group of Inhumans they were tossed into the ravine and were forced to feed off the corpses, infecting their bodies with the highly invasive Skrull DNA which mutated them and caused them to endure for centuries[18]. In the year 1776, A Skrull arrived on Earth and briefly disguised himself as Thomas Jefferson during the writing of the Declaration of Independence and the Bill of Rights. The Skrull attempted to to affect the writing of the Bill of Rights so that slavery was still allowed in early America so that humans would be prepared for a life of slavery during an eventual Skrull invasion. However this plot was thwarted by the Fantastic Four heroes from the 21st century (Who in future years had been the Skrulls longest enemies) who were searching for a Chronal-Anchor. The FF foiled the plot and their leader Mr. Fantastic used hypnosis to turn the Skrulls into cows, a trick he used on the Skrulls that he used on the aliens the first time he met them a number of centuries in the future.[19]

The next recorded incident of Skrulls active on Earth occurred in the 19th century during the days of the American Frontier. A Skrull invasion party set up operations in the town of Wilcox, Oklahoma. The Skrulls there disguised themselves as famous outlaws in order to steer people away, however the local Apache tribe became aware of their duplicitous nature if not their origins. A chance encounter with outlaws Kid Colt and Arizona Annie uncovered the truth and the two gunslingers fought for their lives, decimating the entire town with dynamite, slaying the Skrulls who lived there[20]. The Skrulls were not reportedly seen on Earth until the early half of the 20th century. By this point, Skrulls living on the planet Kral IV established Slave-Masters who took to abducting aliens from other worlds. They kidnapped mobster Machine Gun Martin as he had escaped from prison. The Skrulls on Kral became highly interested in the styles, mannerisms and society of the prohibition era America through their interactions with Martin, many Skrulls on the planet adopted human forms for formed gangs and adopting the culture. Skrull mobsters such as Boss Barker, Lippy Louie, and others resolved mob disputes with gladiatorial combat between their slaves.[21]

Velmax infiltrates the United States government circa the 1940s and 50s.

A Skrull scientist at one point stated that he was the earliest attempt at a "super" Skrull, being experimented on in the 1930s. The resulting experiment did not endow him with super-powers but instead a leg encased in stone, which rendered him useless as a soldier.[22] In 1947, the last known prime Skrull survivor attempted to flee capture from the Deviant Skrulls and his ship crash landed on Earth in Roswell, New Mexico. He was quickly captured by t the United States government who experimented on him for years.[23] Another ship, part of an scouting party for an upcoming invasion also crashed on Earth in the American southwest during that year. aboard the ship were Skrull agents Velmax and Zuhn. Exiting their ship Zuhn was shot by a hunter named Jacob Scott who died in a car crash trying to flee the scene. Velmex stole Scott's identity and fled the scene, leaving Zuhn and their ship to be recovered by the government. Velmex used his identity to work his way into the ranks of the CIA to learn what he could about the Earth's defensive capabilities, but soon came to love the Earth and it's culture and decided to thwart his master's plans[24]. In the early 1950s, Velmax considered taking up the role of Captain America by decided against it[25]. Velmex was not the only Skrull active on Earth at that time. In 1958, a Skrull spy named Zirksu who came on a infiltration mission. He kidnapped test pilot Chuck Chandler and tried to probe him for answers. Chandler managed to escape and bombarded with strange energy he was transformed into the 3-D Man and imprinted on his brother Hal's glasses, but could be freed at will. Zirksu meanwhile attempted to subvert American society, taking advantage of the Cold War hysteria at the time and posing as Diabolik, a communist agent. However this initial attempt at causing hysteria in America was thwarted by 3-D Man, but Zirksu escaped[26].

Zirksu then changed his motus operandi, deciding to use pop-culture as his weapon to further the Skrulls invasion plans. As the leader of a biker gang he attempted to eliminate any sighting reports of his invasion fleet[27]. Posing as recording studio agent Doc Rock, he attempted to manipulate American youth through the music of up-and-coming rock 'n' roller Vance Rivers[28]. Both plots were foiled by 3-D Man. After failing to free Rivers from prison, Zirksu took on the form of then Vice-President Richard Nixon and assisted in empowering Eric Sinkovitz, who attacked 3-D Man as the communist themed Cold Warrior. When 3-D Man defeated Cold Warrior, Zirksu abandoned his Nixon disguise and has not been seen since[28]. Also in 1958, Velmex tracked his ship to a government facility in Long Island, New York. There he encountered the super-hero team known as the First Line and prevented the ship from falling into communist hands. Exposed to the energies of his ship, Velmax briefly reverted back to his Skrull form, but convinced the heroes that the energies mutated him into a shape-shifter and joined the First Line's ranks. Unknown to Velmax, his commander Zuhn survived the past decade and escaped the facility to resume his mission[29].

By November 1963, Zuhn had aligned himself with super-powered villains of Earth under the guise of Chimera and attacked a Stark Industries facility in Dallas Texas on the 21st of that year with the super-human known as Lupine. The First Line stopped Chimera from stealing technology he needed to get off planet, and he was slain in the process, although Velmax kept Chimera's true nature a secret from his teammates[24]. Later on July 16, 1969, Skrull invaders attempted to sabotage Apollo 11 moon landing using Earth operatives Lupine, Typhoon, Positron, Blackjack, and Axis, but this attempt was also thwarted by the First Line[30]. With a Skrull invasion impending an President Richard Nixon threatening to dismantle the First Line, Velmax used his inside knowledge to expose the Watergate Scandal, which ended in Nixon's impeachment, and allowing the First Line to continue to operate—if only underground.[31]

The Skrull Beatles invade the music scene.

On top of their on going invasion plans, the Skrulls sent yet more spies to infiltrate Earth society. A group of Skrulls apparently replaced the popular British rock group the Beatles sometime around what music critics called "The British Invasion" with the intention of infiltrating Earth through popular culture. Another group apparently took the places of the group known as Gerry and the Peacemakers as well. However, like the Skrulls of Kral IV, these Skrulls soon came to adopt the culture and continued their music careers. By December 1974 the "Skrull Beatles" went their separate ways, "Skrull John" left to join Captain Boko in the Kree Liberation Army, "Skrull George" going on a pilgrimage into the Dark Dimension with its ruler the dread Dormammu, "SkrullRingo" sought fame and fortune, while "Skrull Paul" went to see how the Skrull versions of Gerry and the Peacemakers were coming along with their invasion plans[32]. While the Skrull Beatles have since resurfaced the Skrull Peacemakers fate is unknown.

Dorrek VII

By this time the Skrull throne was taken by Dorrek VII who slew his predecessor and married his daughter R'Klll. Their coupling gave them a daughter they named Anelle a born pacifist who sought peace, unlike her warmongering father[33]. Dorrek's brother Prince Dezan a Skrull who sought a peaceful way of life began conspiring with the like minded Zabyk and Myrn. When Prince Dezan's peaceful goals were revealed, Dorrek encased him in an iron mask and had him imprisoned. Zabyk and Myrn pushed their goals underground and continued to operate in Skrull society to further their ends[34]. Shortly after his own rise to power, Dorrek became the target of an assassin named Krimonn. However Dorrek managed to thwart this attempt and as punishment entrapped Krimonn in a device commonly known as a Power Prism[35]. Dorrek sent a Skrull named Zankor was sent to Earth with memory implants that left him to believe that he was merely a human, however his ship crashed causing massive head injuries that submerged his memories so deep that he believed himself to be a human aware of some alien invasion, and frantically tried to warn the authorities[36]. However, the truth was revealed when Zankor was directed to young scientist Reed Richards who deduced that Zankor was really the alien as he was the only one able to decipher the language from the recording device that held his proof and he was turned over to the authorities[37].[38] Also at this time, the Skrulls managed to find a weak link in the First Line. When First Line member the Yeti was reeling over the death of his teammate and love Rapunzel, they sent a female operative to gain his trust[39]. Ultimately, the Skrulls pushed forward with their invasion plans, and the First Line along with a squad of their allies and enemies opposed them. Despite betrayal from Yeti, and large casualties, the First Line destroyed the Skrull invasion fleet, sacrificing nearly their entire roster in the process to stop it[40]. Meanwhile, Yeti and his Skrullian lover fled to the Himalayas mountains, where she eventually died[39].

Modern Age

Early Incursions

The first Skrull invaders of the modern age

The Skrulls apparently did not make another attempt to take over Earth until over 10 years ago, shortly after the birth of the Fantastic Four, a new group of super-heroes who ironically had Reed Richards leading the group. Seeing the Four as the only threat to their plans. They sent four of their agents to impersonate the Fantastic Four, committing crimes to discredit the heroes. However the Four managed to escape captivity and uncover their doppelgängers. Richards then forced the Skrulls to take him to their invasion ship and convince the crew that Earth was protected by invincible warriors and giant monsters, using pictures clipped out of comic books. The Skrulls were easily duped and while one of the spies was allowed to return home with the mothership, the remaining three captive Skrulls were forced to shapeshift into cows and were hypnotized into believing that was just what they were[41]. Soon discovering this deception, Dorrek was furious and invested the entire Skrull Empire's fortune in trying to create an artificial means of giving super-human abilities. Their first tests were on a Skrull named Titannus a weakling Skrull unable to shape-shift. He was given enhanced strength and eventually went on a rampage across the Skrull homeworld and eventually left the Skrull planet to find conquest elsewhere.[42] Dorrek's ambitious plot eventually bore fruit and soon a Skrull warrior named K'lrt was endowed with all the powers of the Fantastic Four, however the source of his power needed to be beamed to him from the Skrull homeworld. As the so-called Super-Skrull, K'lrt clashed with the Fantastic Four on Earth until Reed Richards discovered the source of the Super-Skrull's power and used a jamming device to cut off the energy source. K'lrt was then buried in a dormant volcano[43].

Super-Skrull arrives on Earth

However, Dorrek had his scientist eventually increase the power to free the Super-Skrull. Seeking revenge against the Fantastic Four, the Super-Skrull abducted Franklin Storm the father of the FF's Invisible Girl and Human Torch. Using his shape-shifting powers, the Super-Skrull took Franklin's place in prison, where he had been serving time for vehicular homicide. Using his powers to break out of prison, the Super-Skrull developed the identity of the Invisible Man to commit crimes and discredit the Fantastic Four in the public eye by making it appear that they were letting the Invincible Man get away with his crimes due to his "family connection" to the Fantastic Four. However, Richards eventually deduced the true identity of their foe and built a weapon that threatened to destroy the Skrull throneworld unless they returned Franklin and took the Super-Skrull away. Dorrek agreed to these terms[44] however his warlord Morrat placed a bomb on Franklin to try and kill the Fantastic Four in a double cross in order to curry Dorrek's favour and win the right to marry Anelle[45]. However this plan was thwarted when Franklin sacrificed his life to save those of his children[44]. For his failure to defeat the Fantastic Four, the Super-Skrull was punished and forced to carry out menial tasks for the empire as punishment[46].


Following Franklin's death, the Fantastic Four travelled to the Skrull throneworld to avenge his death. There they clashed with Warlord Morrat, and were easily captured due to their powers failing them on the Skrull planet. By this time, Morrat had grown impatient and sought to use the capture of the Fantastic Four as leverage to oust Dorrek as ruler of the Skrulls. However Anelle informed her father and he ordered Morrat's execution. The Four managed to break free and restore their powers. When Dorrek's firing squad shot at Morrat, Anelle tried to shield him, but she was in turn shielded by the Invisible Girl and Morrat was killed. In thanks for protecting his daughter, Dorrek allowed the Fantastic Four to return home, offering them the false promise that the Skrulls would no longer attack the Earth.[45] Also around this time the Krellek Council assigned five young Skrulls to impersonate and discredit the young mutant heroes known as the X-Men. They did so by unleashing some of the Mole Man's creatures on the surface. However they grew sympathetic to the X-Men and assisted the group in subduing the creatures before they were reigned in by their superiors.[47] The Super-Skrull was later briefly manipulated by the Fantastic Four's enemy Dr. Doom who used his Emotion Changer device to set a number of super-villains to crash the wedding of Reed Richards and Sue Storm. In the end Super-Skrull and the other attackers were shunted back to where they belonged with no memory of the attack when Reed was given a Time Displacement device by Uatu the Watcher[48]. When the Skrull's detected the approach of the planet eater Galactus, they used a device to shield their world from his view, directing Galactus to the planet Earth[49], however he was ultimately defeated by the Fantastic Four.[50] About this time, the Super-Skrull was manipulated by the Asgardian trickster god Loki into attacking his half-brother Thor, but was soundly defeated[46]. Yet another Skrull infiltrated Earth society, this time within the FBI, an agent named Baker was replaced by a Skrull. "Agent Baker" then briefly replaced X-Men liaison Fred Duncan. Baker had the X-Men's Beast investigate recent reports of alien sightings before being transferred to another department.[51]

Skrull mobsters of Kral IV

The Skrull crime bosses on Kral IV later sent their Slave-Master to Earth to capture the Thing, the strongest member of the Fantastic Four[52]. The Skrull succeeded in doing so by disguising himself as Reed Richards and catching the Thing off guard and abducting him[53]. Soon, the Thing arrived on Kral IV, where he was forced into the gladiatorial arenas as a slave of Boss Baker and made to train to fight against Lippy Louie's champion the Mekkan warrior Torgo[21][54]. However his comrades in the Fantastic Four tracked the Thing down and brought down the Skrull Slave-Master and the Skrull mobsters, freeing Ben and starting a revolution from the slaves, with Torgo as their leader.[55]

Emperor Dorrek VII observes Kree activity on Earth

Resumption of Kree/Skrull War

With the awakening of Kree Sentry #459[56] the Kree empire renewed it's interest in the planet Earth, sending a crew on an information gathering mission, among the crew was decorated Kree soldier Captain Mar-Vell, whose early interactions on Earth were mistaken as a new super-hero calling himself Captain Marvel[57]. The increase in Kree activity on Earth prompted Emperor Dorrek to send the Super-Skrull to learn what the Kree were up to. On Earth, the Super-Skrull clashed with Mar-Vell near Cape Canaveral, Florida[58]. However before the Super-Skrull could probe Mar-Vell's mind and learn his mission, he fell victim to his own hypnotic gaze and was commanded by the Kree soldier to flee the Earth[59]. Not long after this a Skrull device crash landed near Cape Canaveral that was purposely sent there to scan DNA information of Earth warriors. It was collected by Carol Danvers and Mar-Vell, in his alter ego of scientist Walter Lawson. When Danvers touched the device the Skrulls copied her DNA allowing them to assume her form almost perfectly. A Skrull invader then posed as Carol and attempted to kidnap Mar-Vell in an Earth space shuttle launch, but he was thwarted by Danvers who rescued Mar-Vell.[60]

It was around this time that the cosmic entity known as the Grandmaster came into possession of the Power Prism which still had Krimmon imprisoned inside. This device was given to transform Ugandan finance minister Kinji Obatu into Doctor Spectrum, part of the Squadron Sinister a group of powered villains that the Grandmaster created to pit against the heroic Avengers who were pawns of the time traveller Kang in a contest[61]. The Avengers defeated the Squadron but Obatu retained the power prism[62][63].

Renewed hostilities in the Kree/Skrull War.

Meanwhile, the Kree targeted Earth again with their then leader Ronan the Accuser attempting to devolve the human race into primitive cave men, leading to a clash with Captain Marvel and the Avengers. However Ronan abandoned this plot when the Skrulls resumed full on hostility against his homeworld[64]. It was about this time that the the lone Skrull that escaped the Fantastic Four in their first encounter with the Skrulls began working with the Super-Skrull who had been exiled following his last failure. The pair used a hyper-beam to revive the Skrulls on Earth who were hypnotized into thinking they were cows[65]. They then joined the Super-Skrull in a plan that K'lrt hoped would win him the favour of his Emperor and win the hand of Anelle in marriage in a bid to gain control of the empire[17]. When the Avenger's battle with Ronan became public knowledge[66] one of the Skrull spies took the place of US Presidential Candidate H. Warren Craddock[67] using his political position to spread xenophobia toward the Kree in general, Mar-Vell in particular and to bismerch the reputation of the Avengers. He also formed the Commission of Alien affairs to begin witch hunts to find aliens hiding among the population. When the Avengers helped Mar-Vell go into hiding this turned the public against them. They were also brought in for questioning by the commission under "Haddock". Meanwhile, the Skrulls captured Mar-Vell and his confidant Carol Danvers as their safe house. The remaining three Skrulls then posed as founding Avengers Captain America, Thor and Iron Man,

Former Skrull Cows posing as the Fantastic Four.

and ordered the current roster to disband[66]. Going to reconvene with Mar-Vell, the Avengers were attacked by the three Skrulls, who then captured the Scarlet Witch and her brother Quicksilver, but the Vision escaped. He brought the real founding members of the Avengers with him later, while the Super-Skrull—posing as Carol Danvers—tried to trick Mar-Vell into building a Omni-Wave Projector a communications device which could also double as a devastating weapon. Mar-Vell saw through the ploy, but this did not stop K'lrt from knocking him out and escaping the arriving Avengers, who bested the three Skrull spies[65]. The three captured Skrulls were once more forced to resume thinking they were cows, where they eventually ended up in the town of King's Crossing[68]. While the Super-Skrull took Mar-Vell and the prisoners to the Great Refuge of the Inhumans, whom he knew were empowered by the Kree and tried to destroy it, but the city was too well protected. He then took his prisoners to the Andromeda galaxy to deliver to Emperor Dorrek, the Avengers were attacked by "Haddock" using SHEILD Mandroids. While on Tarnax IV where Dorrek subdued the Super-Skrull and he forced Mar-Vell to agree to build an Omni-Wave Projector in exchange for the Scarlet Witch and Quicksilver's lives[17]. Back on Earth the Avengers fought free of the Mandroids and assisted Inhuman leader Black Bolt from liberating his people from his mad brother Maximus, but the Avenger's ally Rick Jones was captured by the Kree[69].

Skrulls immobilized by the Destiny Force.

Hostilities by this point had escalated to the point where Dorrek ordered a Skrull armada to destroy the Earth. The Avengers, on their way to rescue their friends intercepted the invasion party and then fought them head on. Mar-Vell meanwhile tricked Dorrek into thinking his was compliant but used the distraction to break free. While on the Kree world of Hala, Rick was imprisoned with the disposed Kree ruler, the Supreme Intelligence who awoken the dormant Destiny Force within Rick[69]. With the Intelligence's guidance, Rick used the Destiny Force to immobilize the Kree and Skrull armies, while the Avengers halted the Skrull's attempt to blow up the Earth, while on Earth the Skrull posing as H. Warren Craddock was reverted to his Skrullian form and beaten to death by an angry mob[67]. In the aftermath of the battle a Skrull Monsteroid crash landed on Earth and was briefly used by both the Puppet Master[70] and Master Khan[71]. The robot was stopped by Spider-Man, the Vision[70] and Iron Fist[71] respectively. With the Kree-Skrull War over for the time being, Captain Mar-Vell met and conspired with Anelle to try to bring peace to their worlds. While a solution was not found, a brief romance between the two impregnated Anelle with a child[72]. When born, Anelle named him Dorrek VIII. Learning of his grandson's mixed heritage, Emperor Dorrek ordered the baby killed, but Anelle managed to smuggle the child to Earth where he spent the coming years unaware of his royal heritage[73].

Skrulls experimenting on Reed Richards following the capture of the Illuminati.

Also as a result of the war, Reed Richards, Iron Man, Black Bolt, Sub-Mariner, Professor X, and Dr. Strange form the Illuminati a clandestined group that would monitor and respond on super-human incidents. Their first order of business was to travel to the Skrull throneworld where Dorrek VII was being appraised of their losses in the war, primarily due to Earth's super-human population. The Illuminati provided a stern warning to Dorrek, telling him to no longer invade the Earth or suffer the consequence, and demonstrated this by blowing up a Skrull warship. However, the Skrulls managed to capture the humans and put them through torturous experiments to learn the secrets of their abilities, under Dorrek's orders. The heroes eventually broke out and managed to flee the Skrull homeworld and return to Earth. In the aftermath, Dorrek learned that they completed collecting all the data they knew on human mutation, magic, and technology and ordered his Priests of Science to get to work engineering Skrulls who could mimic all those skills, no matter how long it would take. Dorrek was intent to fullfill the Skrull prophecies that foretold that the Skrull homeworld would be destroyed and that Earth would be their to take.[74] Following the attack, Dorrek was confronted by Dard'van priestess Veranke who demanded that Dorrek listen to the prophecies of their religion, warning him that the Skrull Empire was doomed and that Earth was their destiny.

Skrulls torturing a Reed Richards clone for answers

Not only did Dorrek dismiss these prophetic warnings, he had Veranke banished to the now deserted Skrullos where she remained in exile for years. The Skrulls first attempts at replicating their human foes came in the form of Reed Richards, whom Dorrek promptly shot in the head, disgusted by the fact that his scientists used the man who turned one of his relatives into a cow[75]. Experiments continued to take place as the Skrulls attempted to find a means of evading notice of the Illuminati by creating clones and having them act out scenarios. Dorrek was frustrated by the constant failures until one Skrull scientist named Dro'ge suggested merely cloning Reed Richards and creating scenarios to trick him into using his intelligence to come up with the ideal way of creating a race of Skrulls that were immune to detection. After using threats and violence towards the clone's "family", Dro'ge came up with a simpler solution, using Richard's love of his son Franklin Richards and using a Skrull to pose as the boy to trick the clone into puzzling out the solution in order to allay his "son's" fears of possible Skrull infiltration. After coming up with the solution on paper, the Richards clone was killed and Dro'ge began to work on making the technology work.[76]

Recovery from the War

The Skrulls were slow to recover from the outcome of the Kree/Skrull War. A number of Skrulls ended up aligning themselves with the mad Titan Thanos, including the Super-Skrull and Skragg. Thanos sought to obtain a Cosmic Cube that was created on Earth and targeted Captain Marvel and his partner Rick Jones to try and obtain it's location. After a failed attempt by Skragg to drive Mar-Vell and Jones mad[77], they attempted a physical attack by tricking him into fighting the Thing, this also failed and Skragg was slain by Thanos. Ultimately, Thanos stepped in and captured Mar-Vell and Jones[78]. Super-Skrull accompanied him to the moon of Titan where Thanos extracted the location of the Cube and left Super-Skrull in charge. Mar-Vell was freed by Mentor and Starfox of the Eternals of Titan, they defeated Super-Skrull in combat and retook the moon for the Eternals. The Super-Skrull meanwhile was left for dead[79]. Eventually, Thanos gathered an invasion fleet to attack the Earth, many of his troops were Skrulls who were slain when the Avengers thwarted the invasion[80]. The Super-Skrull managed to flee to Earth where he took on the guise of Joshua Plague and took control of a band of raiders calling themselves the Rat Pack, using them to steal experimental serums to heal his injuries from battling Mar-Vell. The group clashed with the feline hero Tigra[81] when they slaughtered a Cheyenne reservation to obtain a mystical object called the Soul-Catcher[81]. This slaughter also brought the hero Red Wolf on their trail as well[82]. The two heroes tracked down the Rat Pack and in the heat of the battle, the Super-Skrull revealed his true identity to them[83]. Unfortunately for the Super-Skrull his plan backfired when he attempted to use the Soul-Catcher against Tiger and found himself sucked into the object instead, trapping him inside[84]. The Soul-Catcher was then turned over to the Fantastic Four for safe keeping[85].

Medieval Skrulls

Around this period, the Fantastic Four were tasked with trying to save Counter-Earth from being consumed by Galactus. To this end the High Evolutionary sent two teams to different worlds suitable for Galactus to consume to ensure they weren't inhabited[86]. One team consisting of the Human Torch and the New Man called Gor found a world where a group of Skrulls were living out fantasies based on Earth's medieval period. The Skrulls dropped their guise and explained their being there after the Human Torch accepted a challenge to battle a dragon. With the last of the dragons on that world destroyed, the Skrulls had no reason to stay and left that world to find new worlds to master[87]. Ultimately the Fantastic Four won the battle when Galactus was seemingly destroyed[88]. But he would eventually return to consume worlds again. Back on Earth, a battle between Spider-Man and the villain called Equinox at the Baxter Building, headquarters of the Fantastic Four, resulted in the Soul-Catcher being knocked loose[89]. This allowed the Super-Skrull to free himself, and he battled both Spider-Man and the Human Torch[85]. The Skrull managed to escape and sought out a power crystal to build a warp drive capable of allowing him to finally escape from Earth. However this attracted the attention of Ms. Marvel who assisted Spider-Man in pursuing the Super-Skrull. The Skrull was defeated when Spider-Man created a jamming device to prevent the Skrull from gaining more power reacted with the power crystal, opening a warp gate that the Super-Skrull was seemingly lost in forever[90].

The Skrulls later resumed plotting against their age-old enemies the Kree. A lone Skrull infiltrated the Kree empire, posing as Por-Bat a member of the Kree Science Council and instigated preparations for a conflict dubbed the War of Three Galaxies. He then tried to trick the Kree into using the Inhumans as soldiers on a supposed "sneak attack" on the Skrulls. However the Skrulls had laid a trap and were prepared to destroy their enemies once and for all. The plan was foiled however when the Science Council was interrupted by Captain Marvel and Black Bolt who held off their opening strike long enough for Mar-Vell to reveal that Por-Bat was really a Skrull spy. The Kree put a halt to their attack plans for the time being[91]. Shortly after this, former ruler of Kral IV, Boss Barker, decided to get revenge against the Thing for ruining his dominion over the planet. To this end, he be-headed the robot Torgo and went after the Thing on Earth. Captain Marvel detected Boss Barker's presence on Earth and came to the Thing's aid. In the end it was the severed head of Torgo who put Barker's threat to an end, when he used the last of his energy to slay Barker before he could finish off the two heroes[92].

The Death of Dorrek VII

Skrull invasion of Xandar

Emperor Dorrek then turned his sights to the planet Xandar one of the last worlds in the Andromeda Galaxy that was not already under Skrull rule. The Xandarians sent their ruler Adora to Earth to seek the help of the Fantastic Four. A Skrull assassin (really a robot) followed her, but it was waylayed and left for dead by the FF. Mr. Fantastic, Invisible Girl and the Thing agreed to help Adora defend her world[93]. As the heroes transported to Xandar[94] the Skrull assassin—having survived—hide himself in the Baxter Building to await the team's return[95]. When the Fantastic Four arrived on Xandar, they drove back the latest wave of Skrull attackers. Emperor Dorrek then became aware of another ship heading to the planet and ordered his troops to destroy it[94]. This ship was formally owned by the Nova Corpsman Rhomann Dey and was enroute to Xandar from Earth. It happened to be carrying the hero Nova, his allies Powerhouse, the Comet, Crime-Buster and their foes the Sphinx, Dr. Sun, and Diamond Head. This combination of super-humans easily fought off the Skrull armada and continued toward Xandar[96]. Meanwhile the Fantastic Four continued to battle the Skrulls on the planet, but were abducted and brought to the Skrull homeworld[94]. There Emperor Dorrek had them put before a trail by the Jury of Six who sentenced them to death for their "crimes" against the Skrull empire. The three members of the team were then blasted with an ageing ray by the Skrull executioner Jaketch. As the FF broke free and attempted to make their escape, Dorrek's estranged queen R'Klll returned to oversee her husbands attack on Xandar and found that Dorrek was a coward and began aspiring to take over his rule[33]. The Four were rescued from space by Nova and his party and brought back to Xandar and they brought the Human Torch to the planet with them. However the Sphinx absorbed information from the Xandarian Worldmind and fled to Earth. Meanwhile Protector of Xandar led warriors to fight the Skrulls in space[97]. The Fantastic Four then left to find Galactus to help stop the Sphinx, leaving Nova and his newly formed New Champions to defend Xandar from further Skrull attacks. While on the Skrull homeworld, Dorrek had decided to cut his losses but before he could pull his troops back he was murdered by R'Klll, who then took command of the Skrull Empire. He first order was to launch an all out attack on Xandar[98].

Death of Dorrek VII

Back on Earth, the Fantastic Four brought Galactus with them to deal with the Sphinx as the effects of the Skrull ageing ray began to reach its fatal conclusion. They were unaware that the Skrull they felled earlier was still alive and plotting against them[95]. After both the Sphinx and Galactus were defeated, the ageing Fantastic Four were forced to be put in stasis with the Human Torch the only member unaffected[99]. The robot Skrull assassin then attacked Johnny while his guard was down, but the Torch destroyed the robot. He revived Reed who used the robots technology to build a device that would reverse the effect of the ageing ray, restoring the stricken Fantastic Four to normal. Meanwhile, the Skrulls continued to launch attacks against Xandar[100]. Another Skrull attempted to manipulate the Avengers into going back in time and collecting the Resurrection Stone for him. However the Avenger known as the Beast smashed the stone and the Skrull was ultimately defeated[101]. A group of Skrulls later attempted to assassinate Reed Richards by crashing their ship into the Baxter Building, but their attempt was stopped by the super-powerful Sentry[102]. Also around this time the Super-Skrull kidnapped both Wolverine and novice X-Man Kitty Pryde taking them into space to examine as part of an invasion plan.[103] The pair were assisted by the arrival of Captain Marvel, who helped the two mutants escape to Earth and foil Kl'rt's plans.[104]

Empress R'Klll

One of her first duties as Empress to the Skrull Empire, R'Klll assigned a Skrull named Raksor to act as an ambassador to the Shi'ar who at the time were seeking to persecute Phoenix of the X-Men for the destruction of the D'bari star system[105]. Raksor accompanied the Shi'ar along with Bel-Dann his Kree counterpart to the Blue Area of the Moon where the X-Men were to battle the Shi'ar's Imperial Guard to determine the fate of the Phoenix. Raksor was unimpressed to work along side a member of the Kree. When Raksor tried to attack the X-Man Wolverine, he was rescued by Bel-Dann. Furious at being saved by a hated enemy the two began to fight among each other deep below the surface of the moon[106]. They would spend the next few months below the surface of the moon fighting, completely oblivious to events that would later unfold[107].

Empress R'Klll

Out in deep space, the Skrulls began working on a matter transference device to boost their war effort. They found the key when their device linked to Reed Richard's similarly designed energy transfer device on Earth. Seeking to obtain Reed's device, and fearing interference from the Kree, the Skrulls sent agents to stop Captain Marvel while sending the best warriors—dubbed the Prime Ten—to Earth. The Prime Ten ambushed the Fantastic Four at their headquarters, but were ultimately defeated by the Four and Mar-Vell. Using the transference device they sent the Skrulls back to their asteroid base causing the device to overload and explode, ruining their plans of ever using it against the Kree[108]. While back out in Skrull space, the Skrulls managed to defeat Protector and his warriors thanks to the assistance of Diamondhead who betrayed his "allies", slaying Crime-Buster in the process. Shunting Protector and his crew into another dimension a Skrull war party disguised themselves as the crew and returned to Xandar to launch an ambush attack. There they were opposed by Nova, Powerhouse and the Spaceknight Rom who was on Xandar trying to learn the location of his homeworld Galador. With the assistance of Rom, the New Champions managed to thwart the Skrull invasion[109].

When it was learned that Captain Marvel was dying of cancer, the Skrulls sent General Zedrao to award their greatest enemy a Skrull medal of valor and pay his respects on behalf of the Skrull Empire[110]. However the duplicitous Skrulls also took advantage of this situation to continue their constant infiltration and spying. Among the gathered heroes come to pay their respects to the dying hero was a Skrull named Bartak who initially infiltrated the gathering posing as the hero known as Devil-Slayer. However upon noticing that the real Devil-Slayer was in attendance, Bartak changed his shape to resemble the Silver Surfer who at the time was exiled and stranded on Earth after betraying his former master Galactus. When the heroes inquired as to how the "Surfer" managed to escape the Earth, Bartak convinced them that Galactus had set him free. His deception a success, Bartak returned to the Skrull homeworld and underwent augmentation to imitate the Surfer's powers and continued to pose as the hero in space while he was still exiled on Earth.[111] The Skrulls soon planned another invasion of Earth, the plans of which were learned by the Shi'ar who sent their agent Gladiator to warn the Fantastic Four. However Skrulls impersonating the X-Men intercepted Gladiator and convinced him that the Fantastic Four had been replaced by Skrulls leading to a battle between the famed group and Gladiator[112]. However with the assistance of Spider-Man and Captain America, the Fantastic Four managed to reveal the Skrull trick and defeated the X-Men imitators. They were then taken into custody of Gladiator who took them into space[113].

During all this time, the Skrulls who were trapped in the form of cows on Earth were being milked in the town of King's Crossing. When the towns people began ingesting the milk their bodies were infected with the invasive Skrull DNA. These now half-Skrull half-humans became increasingly xenophobic and began shutting off the town while mass producing the Skrull DNA infused milk. When the Fantastic Four's friend Sharon Selleck stumbled upon the town and was forced to stay, she managed to contact the Fantastic Four who came to her rescue. Getting a sample of the tainted milk, Reed Richards managed to develop an antidote and cure the locals and destroy all the tainted milk and crops that were irrigated with it[68]. The Skrull cows however where not located at this time and continued to endure for a time[114], some of them even began breeding producing Skrull/Cow hybrids.[115] R'Klll was also in talks with the Kree Supreme Intelligence and they made an agreement that the outcome of the Kree/Skrull War would be determined by proxies and selected both Raksor and Bel-Dann on the moon. They had the Watcher gather the two combatants and they agreed that whoever one the one-on-one battle would then determine whose race was the victor in the war[107]. Also around this time a Skrull scholar named Klobok was exiled from the Skrull throneworld for his obsessive research of magic.[116]

Destruction of the Skrull Throneworld

Destruction of Tarnax IV

The Skrull homeworld soon came under threat of consumption by Galactus when his newest herald Nova who was able to detect the Skrull throneworld even though it was cloaked, as in previous close calls with the world devourer[117]. Shortly before the arrival of Galactus, a group of X-Men from the near future found themselves transported to the Skrull homeworld by the other-dimension being known as Ejulp[118]. There the group discovered an elaborate training facility that was designed to resemble New York City where Skrulls were training as disguised super-heroes for yet another planned invasion of Earth. These new Warskrulls were designed to be locked into the form of those they impersonated. The X-Men clashed with their spies-in-training who posed as various super-heroes both dead and alive, including some based after the X-Men themselves.[119] The battle raged until suddenly Galactus arrive to consume the planet[120]. The X-Men's leader Professor X then attempted to convince Galactus not to consume the Skrull throne world, however his efforts were in vein. The X-Men, were then forced to flee the planet in a ship that would pilot them back to Earth, on an extended course that would bring them home in their proper time.[121] Despite their efforts to fight off Galactus, the world devourer succeeded in consuming the Skrull throneworld. Among those slain in the destruction were Empress R'Klll and her daughter Anelle.[122] Reed Richards was ultimately blamed for the destruction of the Skrull throneworld due to the fact that he previously saved the life of Galactus.[123] He was then kidnapped by the Shi'ar who were acting on behalf of the Skrulls at the behest of Prelate Karant Kiar.[124] During the so-call "Trial of Galactus", Karant Kiar sought to have Richards found guilty for the slaughter of his homeworld and executed. However testimony from the Watcher, Asgardian All-Father Odin, the embodiment of the universe known as Eternity and from Galactus himself convinced the court that Galactus was an integral part of the universe and the destruction of the Skrull homeworld was part of a greater destiny, and so Richards was acquitted of all charges and released[124]. Following the destruction of their homeworld, Skrulls that were determined to have mutant abilities were no longer culled, and were used for experimentation and weaponization purposes, although they passed along the legend of Charles Xavier, the man who tried to save the throneworld when none of the Skrull would stand up to the world devourer.[125]

One of the few survivors of the destruction of Throneworld was the son of the Skrull scientist who helped created the Super-Skrull. When Galactus was destroying throneworld, he used his technology to endow his son with enhanced abilities and sent the boy to Earth in the hopes that he would conquer the Earth[126]. However when this Skrull child's ship landed in the American mid-west, he was found by a devoutly religious family, who unaware of his Skrull heritage raised the boy as their own under their Christian values system under the name Ethan Edwards[127].

Skrull Civil War

With the death of R'Klll five[128] then six Skrull governors all vied for control of the empire. These governors included Gorath,[129] Kylor,[130] Yorak,[131] an unnamed overweight female Skrull,[132] S'Byll of Satriani,[133] and Karza.[111] All began their own agendas and the Skrull empire fell into complete chaos, with the governors of each Skrull world claiming the title of emperor or empress. Civil war broke out between the various Skrull worlds claiming more lives than Galactus destroyed in his consumption. This made the Andromeda galaxy ripe for the picking for the space pirate known as Nebula the self-proclaimed daughter of Thanos the Mad Titan and she began raiding world after world, eventually forming an alliance with Skrull governor Gorath[129]. It was during this ensuing chaos that Skrull scientists Myrn and Zybek were reunited, and they began working on a Hyper-Wave Bomb that would change their empire completely. However they were unaware that their entire plan was overheard by Raksor who sought out allies to stop their mad schemes[34].

With their empire crippled, the Skulls began focusing on other external threats. The Dire Wraiths, offshoots of the Skrull race, were gaining a foothold on Earth and other worlds despite the intervention of the Spaceknight Rom. As such the Skrulls sent a landing party to the town of Clairton West Virginia, a town that was a beachhead in the Wraith invasion of Earth. The Skrull's arrival coincided with the Dire Wraith's final push to wipe out the town and defeat Rom and his ally Starshine, whom the Wraiths banished to a shadow dimension[134]. The Skrulls held off the Dire Wraiths until Rom and Starshine could free themselves and assisted the Spaceknights in wiping out the Wraiths there. Needing to return to their crippled empire, the Skrulls left but their leader—named Supremor warned Rom that if he failed to exterminate the remaining Wraiths on Earth, the Skrulls would destroy the planet[5]. Over the past months, the Super-Skrull had been caught in a hyper-space gateway travelling to the Skrull homeworld. However with the destruction of the planet at the hands of Galactus, the Super-Skrull found himself shunted back to Earth[135]. His return to Earth brought him to the Mount Logan Cosmic Research Station in the Yukon territories of Canada. There he decimated the crew and clashed with the Canadian super-hero known as Sasquatch[136]. When it became determined that K'lrt had contracted an advanced form of Leukaemia from the months spent travelling unshielded through hyperspace radiation, the Sasquatch agreed to build a device that would transport the Super-Skrull back to Earth. However the hirsute hero was merely waiting for the Skrull to buckle in pain from his illness again to ram him into the partially completed warp-gate, seemingly destroying the Super-Skrull[135]. By this time the battle on the moon between Raksor and Bel-Dann interrupted the wedding of the Inhumans Black Bolt and Medusa. The Inhuman royal family and the Fantastic Four then attempted to stop the hostilities between the two. However learning from the Watcher why the two aliens were fighting the Earthlings attack the two combatants and forced them to work together. With the Earth heroes pretending to be slain, the Watcher declared the battle won, telling the Skrull and Kree that the outcome was that both races need to strive together and they were then teleported away[107].

At this time General Kalamari of the Skrull high command decided to focus his efforts on destroying the Fantastic Four by placing an impostor into their midsts. With the cost of creating a new Super-Skrull to imitate one of the Four, Paibok suggested that they instead have a Skrull replace one of their closest allies. They selected Alicia Masters a blind woman who was close friends of the Fantastic Four and at the time was dating the Thing. Paibok selected a female Skrull named Lyja as his candidate and she spent months studying to be the perfect imitation of Masters. They then kidnapped the real Alicia and Lyja then took her place[137]. However the plans had to change as the kidnapping of Alicia coincided with the male members of the group being abducted by the omniscient being known as the Beyonder to fight in the so-called Secret Wars[138]. When they returned to Earth, the Thing stayed behind on Battleworld as he could revert back to human form and he was replaced by the She-Hulk[139][140]. Lyja instead changed her focus from the Thing to his partner the Human Torch and spent the ensuing months romancing him[137].[141] Meanwhile the Super-Skrull was pulled out of his exile in the Van Allen radiation belt by Master Khan who was plotting against his old foe Iron Fist. Khan cured the Super-Skrull of his Leukaemia and then had him transform into a young boy whom he dubbed Bobby Wright, Khan imprinted false memories into the Super-Skrull of Bobby Wright that were so thorough it could fool his old foe[142]. For months, Super-Skrull posed as a young orphan who believed he gained super-powers (and the ability to transform into an adult version of himself dubbed Captain Hero). Bobby and his alter-ego of Captain Hero became integral in the lives of Iron Fist and his partner Luke Cage[143][144][145][146].

Hyperwave Bomb

In the Andromeda Galaxy, Nebula began acting on the behalf of Governor Gorath in demanding that General Zedrao commit his military fleet to his cause. When Zedrao refused, she sent her minion Captain Marvel to spy on the base, unaware that she was a member of the Avengers. Nebula then used her ship the Sanctuary II to blow up the Skrull base, but Marvel helped the Skrulls escape and alerted her comrades the Avengers who came to assist along with the former herald of Galactus known as Firelord. While they mobilized to deal with Nebula and her forces, the notorious space pirate slew Gorath in her bid to take over the Andromeda galaxy herself[129]. With the aid of the Avengers the Skrulls fought Nebula's forces, but the battle was not truly won when the Beyonder arrived seeking to join the Avengers to learn how to be a hero. In the heat of the battle, he teleported Nebula and her minions away, suddenly ending the fight[147]. At this time the prison planet Klarr were attacked Raksor who freed the long imprisoned Prince Dezan. They then got passage to the Power Asteroid where the Zybek and Myrn were working on the Hyperwave Bomb. They travelled aboard Kral IV's "Big Casino" starship which was fashioned after a 1930s era Zeppelin and piloted by a Skrull calling himself Floyd Donahue. Along the way they intercepted the Avengers who had pledged to assist General Zedrao in capturing Nebula and sought the Power Asteroid's equipment to try and track her[34]. At that same time a female Skrull governess sought to secure herself as the empress of the Skrull empire by destroying the Fantastic Four. To this end, she had one of her minions disguise himself as the Elan creature known as the Infant Terrible to lure the Fantastic Four to the Skrull world under the false pretence that Elan had been decimated by the Skrull. Reed Richards saw through this plot and defeated the Skrull governess on her homeworld and took them prisoner. Reed and the Fantastic Four then brought their prisoners to the Power Asteroid to dispose of[132].

Meanwhile, Zabyk completed the Hyperwave Bomb just as the Avengers were attacking his fortress. Revealing that the Hyperwave Bomb would erase the Skrulls shape-shifting ability, trapping them in whatever form they were stuck in. Zabyk then slew Myrn and sealed him in what he believed would shield him from the effects of the bomb, which he believed would make him the new ruler of the Skrulls. Despite their efforts, the Avengers and the Fantastic Four were unable to stop Zabyk from unleashing the Hyperwave Bomb. The weapon froze all the Skrulls in whatever form they were in at the moment the bomb went off.[34][132] Many Skrulls who were on espionage missions found themselves trapped in those forms. These included Al'Arok a spy on Zenn-La;[130] Aptak a male Skrull who was spying on the Waziliah in female form; Nenora who posed as a Kree and was infiltrating her way into the Kree Science Council;[131] A Skrull posing as the Elder of the Universe known as the Contemplator;[148] Bartak the Skrull who was posing as the Silver Surfer.[111] Incidentally, the Super-Skrull, who was still under the guise of Bobby Wright, was still able to change from the child form of Bobby Wright and the adult Captain Hero, likely due to the influence of Khan's magic.[146] Following the incident with the Hyperwave Bomb, General Zedrao and his soldiers then destroyed the Sanctuary II before thanking the Avengers for their help and sending them on their way.[149] It was then greatly accepted by the Skrull Empire as a whole to try to keep the loss of their shape changing powers a secret otherwise it may provide their many enemies with an advantage.[150]

On Earth's Master Khan's manipulation reach fruition. "Bobby Wright" was diagnosed with an illness, that when it came to fruition, as Captain Hero—insane from the pain racking his body—beat Iron Fist to death before he seemingly disintegrated[151]. Not only the Super-Skrull's involvement a mystery, but also the fact hat Iron Fist had secretly been replaced by a H'ylthri impostor[152]. The Super Skrull was not dead, but returned to the hyper-space prison where Khan had pulled him[128].

Renewed Hostilities Against the Kree

In order to keep the loss of their shape-shifting abilities a secret, the Skrulls entered into an alliance with the Elders of the Universe for protection.[150] This however was to benefit the Elders as they were also seeking to destroy Galactus and create a new Big Bang in which the Elders would be the new Galacti of the universe that formed.[3] Unknown to the Elders, their ranks were infiltrated by the Skrull posing as the Contemplator.[148] The Elders convinced the Skrulls to capture Galactus' new herald Nova to force Galactus to destroy Kree worlds only. They then sent the Champion to slay the Silver Surfer on Earth. The Surfer defeated the Champion and learned of Nova's capture. Freeing himself from the barrier that had kept him on Earth for years, the Surfer made a deal to rescue Nova in return for Galactus allowing the Surfer to remain free. The Skrulls prison was no match to keep the Silver Surfer from rescuing Nova and returning her to Galactus.[150]

The Skrulls then Ptakr as an emissary to petition Shalla-Bal the ruler of Zenn-La to join the Skrull in their war against the Kree. Shalla-Bal refused to side with the Skrulls, deciding to remain neutral and warned them against attacking her as her planet were now under the protection of the Silver Surfer. After a failed attempt to assassinate the Surfer, Ptakr and his entourage were ejected from Zenn-La.[130] Not long after this the Silver Surfer and the Avengers were involved in a scheme by the Grandmaster to make all the Elders of the Universe immortal, bringing them a step closer to succeeding in their machinations.[153][154] While the Elders once again sought to destroy the Surfer, Ptakr's crew was attacked by Kree soldiers, but the Skrulls committed suicide before they could be boarded and the secret of their lost shape-shifting abilities could be discovered.[155] Following their bid for immortality, the Elders met with the fake Contemplator among their numbers. The meeting was interrupted by the Silver Surfer and his new ally Mantis who managed to escape destruction. Meanwhile, the Supreme Intelligence of the Kree correctly deduced that the Skrulls lost their shape-shifting abilities. This also coincided with the arrival of the Celestial known as Jemiah the Analyser landed on the Skrull world that was under the rule of Kylor.[156] Kylor ordered Jemiah destroyed, but none of the Skrull weapons were powerful enough to destroy the Celestial. Meanwhile, a Kree spy was sent to confirm the Supreme Intelligence's suspicions, but despite his efforts to disguise himself he was discovered and captured by Skrull authorities.[3]

By this time the Skrull spy Nenora had became a member of the Kree Science Council, who were second to only the Supreme Intelligence. When the Intelligence ordered an attack on a Skrull armada that was passing the Kree world Sandomar II. However the attack was a failure as Nenora alerted Kylor and the Skrull armada withdrew. This however tipped off the Supreme Intelligence to the fact that there was a spy in his midst, but Nenora's cover was so deep that the Supreme Intelligence was led to believe his aid Nullet was the Skrull spy. Kylor's renewed aggression toward the Kree started a feud between himself and another would-be Skrull emperor Yorak.[131] At this time, the Elders of the Universe came to believe that the six Soul Gems had the power needed to destroy Galactus and create a new big bang. Having collected five of the six, they sent the Skrull impersonating the Contemplator to the Kree homeworld where the sixth was under the control of the Supreme Intelligence. The Contemplator was denied the Soul Gem, as Supremor required it to maintain his sanity caused by the racial conflict between the pink and blue skinned Kree minds that made up the part of his collective intelligence. When the Silver Surfer came to warn the Supreme Intelligence to stay away from Zenn-La, the Intelligence used the Soul Gem to trap the Surfer inside the Intelligence.[148] However the Surfer broke free and absconded with the Soul Gem, leaving Supremor insane.[157] The Elders then kidnapped Mantis and Shalla-Bal in order to force the Silver Surfer to turn over the Soul Gem. The Surfer agreed, and the Elders then tried to use them to destroy Galactus, but this plan was thwarted by the Surfer and Nova when Nova destroyed the star which the planets the Elders were using in conjunction with the gems to destroy the world devourer. The resulting blackhole swallowed the Soul Gems, as well as the Trader, Possessor and the Astronomer[158], while the Grandmaster, Gardener, Collector, Champion and Runner were consumed by Galactus, the phony Contemplator escaped and Galactus ordered the Surfer and Nova to apprehend him. While on the planet Hala, Nenora had become the Supreme Leader of the Kree. She then decided to take her ambitions beyond the Skrull empires age old war, and slew her former lover Apatk who came to Hala posing as a Wazilian ambassador.[159]

Meanwhile on Earth, a group of Skrull Slavers disguised themselves as a Kentucky fair in order to collect slaves. Their activities coincided with the arrival of Cannonball, Wolfsbane, Richtor and Boom Boom of the New Mutants who were visiting Cannonball's family at the time. When they visited the carnival, the Skrulls attempted to make them slaves as well, but the Skrulls proved no match for the young mutants and were forced to flee into space[160].

While back out in space, the Silver Surfer and Nova with information from Nenora, tracked the Skrull impersonating the Contemplator in the Coal Sack Nebula the home of the real Contemplator. There the fake gained the protection of the space pirate Cap'n Reptyl who briefly managed to capture the Surfer and Nova. During this period he contemplated the Elder known as the Obliterator renewing his sense of purpose after a defeat at the hands of the Surfer.[161] Meanwhile, under orders from governor Karza, Bartak the Skrull posing as the Silver Surfer began attacking Kree outposts to taint the Surfer's reputation.[162] When the Surfer and Nova broke free, Reptyl and his minions retreated back into the Coal Sack. Angered over this defeat, Retyl then seemingly slew the fake Contemplator, consuming his body[163] and leaving his severed head in space to be found by the Surfer and Nova.[133] As Bartak planned, the Kree blamed the real Surfer for the attacks and Ronan the Accuser was sent to eliminate him, but the Surfer bested him in combat. Meanwhile, Nanora now deciding to rule over the Kree Empire, led Kylor and his armada into a Kree trap, leading to their death. With Kylor's death governess S'Byll decided to make her move to become the next Skrull empress and ordered one of her minions to Earth.[133] The Silver Surfer and Nova meanwhile were attacked by Bartak, who defeated the Surfer in combat and trapped Nova. But before he could get away to cause more chaos, he was slain by Ronan the Accuser who upgraded his weapons to be more than a match for the real Surfer. This was apparently Karza's onyl attempts to vie for leadership of the empire as he was never seen or heard from again. Meanwhile, S'Byll's minion arrived on Earth and fired a device into the air before being felled by a pair of hunters[111]

Empress S'Byll

This blast reconstituted the Super-Skrull who had no memory of his experience as "Bobby Wright" and was led to believe that Skrull science cured him of his cancer as well as freed him. His return coincided with the arrival of the Silver Surfer to Earth. The two battled but were captured by the Eternals who sought to map the DNA of the Silver Surfer and the Super-Skrull on behalf of the High Evolutionary. Unwilling to let the Eternals force their own ends, the Surfer and the Super-Skrull worked together to free themselves. The Surfer, telling the Super-Skrull what happened and that because he was the only Skrull left who could shape-shift, convinced him to return to his empire to try and restore to the glory it once was in the hopes this would end the current Kree-Skrull war[128].

The Skrull posing as the Contemplator survived his previous encounter and managed to relocate his severed head to a moon nearby the Coalsack Nebula and awaited Cap'n Reptyl and his crew to land there. Reptyl left his oafish subordinate Clumsy Foulup in charge, and the faux-Contemplator slowly began mentally manipulating him for his own ends. While Nenora continued to intensify her assaults on the Skrulls, but they were soon reunited with the Super-Skrull who had aligned himself with S'Byll and her faction of Skrulls. Meanwhile, Nanora went to Zenn-La to offer her apologies to the Silver Surfer for her empire attacking him previously when Bartak framed him. This visit prompted to Cotati, who had been using their connection to all plants to start hinting to Empress Shalla Bal that Nenora was really a Skrull.[164] Unfortunately for Shalla Bal, Nenora's visit was also to secretly plant listening devices in her home and place a Kree Sentry to capture her if need be.[165]

By this time the Kree empire formed an alliance with the Badoon, causing massive casualties for the Skrulls. S'Byll and the Super-Skrull then tricked the Silver Surfer into defending the Skrull armada from the Badoon, forcing him to ally himself with the Skrulls in order to defend Zenn-La after the attack would be viewed by the Kree as the end of his neutrality in the war. The Skrulls came to the Silver Surfer's aid in fighting off a Kree invasion fleet that was poised to attack Zenn-La. Arriving on the planet's surface, Shalla Bal also accepted the alliance and S'Byll and the Super-Skrull revealed the reason for the entire deception: Duplicating the machine the Eternals used on the Super-Skrull and Silver Surfer, they devised a way to reawaken the shape-changing ability in S'Byll and give the ability to restore that power to the entire empire. The Surfer and the Skrull then pooled their powers to power the device, not only restoring S'Byll's shape shifting abilities but enhancing them to cosmic levels. She then joined the Surfer and Super-Skrull in driving the remaining Kree ships from Zenn-Lavian space.[166] Nenora then led an entire invasion fleet to attack Zenn-La again, forcing the trio to fight a losing battle. That is until the Silver Surfer managed to broker a deal with Nenora: He would leave Zenn-La forever if she agreed to stop attacking his homeworld, and she agreed pulling her forces back. This coincided with the Contemplator impostor moving ahead his plans, manipulating Clumsy Foulup into contacting the Kree and offering them the means of defeating the Skrulls.[167]

Returning to her homeworld of Satriani with the Surfer and Super-Skrull. There she began using her newfound powers to restore the shape shifting abilities of the Skrulls that resided there. This drew the enigmatic Stranger who came to apparently capture S'Byll. However the Stranger returned her when Cap'n Reptyl came to offer his alliance to the Skrulls and offered the Stranger the mutants aboard his crew.[168] Reptyl's addition to the Skrull forces was not without turbulence, when Reptyl started paying too much attention to S'Byll, the jealous Super-Skrull attacked him but was seemingly slain when Reptyl ripped out his throat.[165] The Super-Skrull survived however, his body recovered by the Badoon who attempted to experiment on him. Fully healed, K'lrt broke free from his captors and escaped[169]. This was all part of a ruse on the part of S'Byll, orchestrated with the intended purpose of making the enemies of the Skrulls believe that the real Super-Skrull was dead and that when the K'lrt returned they would be led to believe he was merely an impostor[170]. At some point during the conflict, S'Byll's uncle dubbed Talos the Untamed due to the fact that he has superior strength although being born without a shape-shifting ability was defeated and captured by a Kree war party earning him shame among the empire[171].

Meanwhile on Zenn-La, Shalla Bal deduced the secret of Nenora, but her Kree Sentry capture Shalla before she could tell anyone. And on Hala, Clumsy Foulup set the beginnings of a trap for Cap'n Reptyl distracting the Kree enough for the astral form of the "Contemplator" to gain access to the chamber of Supremor.[165] With Shalla Bal prisoner, Nenora had the human named Midnight augmented into a cyborg and launched another attack against the Skrulls. Meanwhile the "Contemplator" began manipulating the pink skinned Kree to further his goals of taking over the Kree empire. In Skrull space, while the Silver Surfer was detracted fighting Midnight, Clumsy Foulup—who had been made Reptyl's second in command after being smuggled back to the moon where he was left prior to his betrayal—seemingly slew Reptyl and made his escape back to the Kree.[172] With the seeming death of Reptyl, the Skrull armada was decimated by the Kree, leaving only S'Byll and the Surfer the sole survivors. They fought their way to Hala where they gained entrance into the Kree palace thanks to Shalla Bal who was freed by the influence of the Cotati. As Clumsy Foulup was being thanked as a war hero by the Kree, they stormed Nenora's throne room. There, under the witness of the Stranger, S'Byll used her powers to restore Nenora's shape shifting ability to slew her.[173] S'Byll then revealed the truth to the Kree and brokered a truce, failing to convince the Kree to declare peace. After she was taken back to her homeworld by the Silver Surfer, the "Contemplator" attempted to put his plan into fruition: With Foulup seen as a hero, especially to the pink skinned Kree, the "Contemplator" would then pose as a "restored" Supreme Intelligence" and have Foulup back him, making the "Contemplator" the secret ruler of the Kree empire. However, Foulup ended up betraying the fake when the Cotati offered Foulup to be the actual ruler of the Kree. When the "Contemplator" attempted to pull his ruse, the Cotati overpowered him with their superior telepathic abilities, obliterating his mind and making it seem like the Supreme Intelligence endorsed Clumsy Foulup's ascension to leadership.[174]

In an unrelated note, a Skrull was prisoner of Captain Orack's slave ship. He was freed by the alien Beta Ray Bill[175]. A Skrull by the name of Kholdsor, trapped in human form due to the Hyper-Wave Bomb, and not yet restored by Empress S'Byll attempted to obtain a copy of Rick Jones' book "Sidekick" in the hopes of learning the secrets of how Rick used the Destiny Force to win the Kree/Skrull War, this mission was a failure[176].

Covers Exposed

Following the end of the most recent Kree/Skrull War, S'Byll was named the new Skrull empress and she ruled from planet Satriani, slowly restoring the Skrulls natural ability to change shape. Her world was briefly invaded by the Impossible Man and his family who made a stop of Satriani from a vacation on Earth to show the Skrulls their vacation slides, much to S'Byll's irritation[177]. Not long after this the Super-Skrull returned to Satriani to a heroes welcome, but was unhappy with the deception as it worked against his honor. However seeking religious guidance from a statue of Sl'gurt. Finding resolve, the Super-Skrull then tracked down Cap'n Reptyle and seemingly slew him. Returning to S'Byll to inform her of what he did, he was stripped of his rank and told to serve under Commader K'Targh's imperial starfleet until further notice[170]. The Super-Skrull however sought to win back the adoration of his leader and so he and a team of Skrulls then returned to Earth in order to capture Rick Jones and harness the Destiny Force as a weapon for the Skrulls. Learning that Rick was currently on a book tour for his new book "Sidekick" they took over the town of Buckport, Utah, and invited him to a phony book signing. They then kidnapped him and tried to torture him into subservience[178]. This failed, and when Rick's allies the Hulk and Betty Ross came to his aid, a battle broke up. Ultimately the Skrull team—including Super-Skrull—were seemingly slain when the energy core of their ship destabilized and exploded[169].

Despite S'Byll's rule over the Skrull empire there were those who sought to remove her from rule. A rogue Skrull (possibly a rare mutant Skrull)[citation needed] named De'Lila attempted to obtain a Skrull Inorganic Technotroid egg to use against the empress however it ended up crashing on Earth.[179] Following it there, De'Lila subdued the Fantastic Four in hopes of using Reed Richard's technology to locate it. However she was followed to Earth by Skrull warriors who took control of the Mole Man's monsters on Monster Island and used them to search for her, angering the Mole Man for this trespass. Seeking to waylay these monsters from finding her, De'Lila used her telepathic abilities and shape-shifting power to resemble the Invisible Woman to trick Spider-Man, Wolverine, the Hulk, and Ghost Rider into thinking the Fantastic Four were dead and to help her. This New Fantastic Four were sent out to stop the Skrulls[180]. Unable to locate the Technotroid on her own, De'Lila used blackmail and her powers to coerce Reed Richards into helping her, successfully tracking it to Monster Island. However this also brought them before the other Skrulls, Mole Man and his Moloid army, and the New Fantastic Four, who had formed an uneasy alliance after piecing together De'Lila's deceptions[181]. Learning what De'Lila was after it was a race to see who got to the Technotroid before it hatched as the creature was made to biologically imprint itself to whomever it first saw. A pitched battle between all three parties erupted over the creatures until the original Fantastic Four arrived to stop De'Lila. In the end, the Technotroid imprinted itself to one of Mole Man's creatures who took it as a child. De'Lila fell victim to the Ghost Rider's Penance Stare and was driven made and turned over to the Skrulls[179]. Returning to the surface the Skrulls and the two Fantastic Four groups were attacked by the Mole Man now that they were out of his domain, however Spider-Man convinced the Mole Man of the futility of their continued battle and the Skrulls were allowed to take De'Lila back to the Skrull homeworld for punishment[182].

Also on Earth, the Super-Skrull survived his encounter with the Hulk and apparently remembering his time as Bobby Wright sought out a new plan to destroy the Earth. At the time Daniel Rand, the hero known as Iron Fist was still believed to be dead. He sought out Ward Mechum the part controller of Rand-Meanchum the organization that both men owned. He offered Meachum the hand of the most beautiful woman in the universe and a world to rule if he assisted in drilling into using the Nuwali devices in the Savage Land to destroy the Earth. Meachum agreed in return for the taste of power and a beautiful woman. The Super-Skrull then used his hypnotic powers to make Ward's niece Joy follow their cause[183]. Disguising himself as Daniel Rand, the Super-Skrull convinced the world that Rand was still alive[184]. "Iron Fist's" resurrection brought the suspicions of his former lover Misty Knight who confronted the phony Daniel Rand to learn the truth, but the Super-Skrull researched his role well and thorough convinced her of Rand's apparent "resurrection"[185][186].

Around this time the Warskrulls program had made many advancements, allowing Skrulls to copy the shapes, powers, memories and personalities of templates that were caught in their Nexus Amplifier devices[187]. To this end they captured the Starjammers and impersonated their members as well as Charles Xavier, former teacher of the X-Men who was travelling with them at the time. Using Xavier's powers they also took telepathic control of disposed Shi'ar empress Lilandra in a bid to take over the Shi'ar throne from her sister Deathbird[188]. They began a campaign of genocide across the Shi'ar galaxy, wiping out many races such as the P!ndyr of Epsilon Seikosha II[189]. Blaming these atrocities on Deathbird, they managed to remove her from the throne and place Lilandra back in her rightful title, all be it a puppet ruler for the Warskrull agenda. This forced Deathbird and teleporter Lila Cheney to seek out the X-Men's aid in freeing the empire[190][191]. The X-Men initially did not believe Deathbird due to the fact that she did not know the full scope of the Skrull incursion and they sided with "Professor Xavier" and the "Starjammers" in apprehending Deathbird. As key members of the X-Men were slowly being replaced by Warskrulls, the X-Men Gambit and Jubilee stumbled upon part of the truth[192]. Jubilee was captured but Gambit managed to escape. The X-Men arrived on the scene, unaware that their teammate Psylock was replaced as well. However Wolverine detected the truth and slew the Warskrull posing as Xavier. The Skrull posing as Psylock knocked out Wolverine and the rest of the X-Men were teleported away by Lila Cheney and Deathbird to Epsilon Seikosha II. A Warskrull named Prime took the place of Xavier and Wolverine and Jubilee were templated as well. Meanwhile the Warskrulls posing as the Starjammers were sent after the X-Men.[187]. The X-Men learned the truth and defeated the Starjammer impostors and returned to the Shi'ar homeworld. During the ensuing battle the real Charles Xavier freed himself and took down the Warskrulls with his telepathic powers. With the primary Warskrulls defeated the X-Men left the Shi'ar galaxy, leaving Lilandra nad her Imperial Guard to weed out any other Warskrulls that might be operating in the area[188].

When the recently revived Thanos gathered the six Infinity Gems he combined their power into the Infinity Gauntlet and used it to wipe out half the universe as a gift to his lover Death. This included half of the Skrull empire. Unaware of Thanos' involvmed, S'Byll suspected that the Kree were involved and called her people to prepare to go to war again[193]. However it appears that this conflict did not happen as a gathering of surviving heroes eventually defeated Thanos and restored the universe to normal[194].

Around this time on Earth, the Super-Skrull and Ward Meachum were bringing their plans to fruition, meeting with Ka-Zar the ruler of the Savage Land and getting his permission to "mine" in the area[195]. However these activities brought Namor the Sub-Mariner and his niece Namorita to the Savage Land to investigate. Super-Skrull, disguised as Iron Fist easily captured Namor[196] and his allies Namorita, Phoebe Mars, and Carrie Alexander, with the help of Misty Knight who believed that "Rand" was really her former lover[197]. Ka-Zar's wife Shanna the She-Devil rescued Namor and his captives and they launched a counter attack forcing the Super-Skrull to reveal himself[198]. As the Super-Skrull battled Namor, his allies convinced Ward to believe was being betrayed when they had him consider what the "most beautiful woman in the universe" would be interpreted by a Skrull. Meachum then tried to smash the machine but the Super-Skrull burned him alive. However this did not stop Namor from smashing the device and saving the Earth, forcing the Super-Skrull to flee[183]. Not long after the departure of the Super-Skrull another Skrull calling himself Samuel J. Skrull was sent to Earth as part of yet another invasion. His ship crashed in the Savage Land and he was worshipped as a god by the natives. When the Impossible Man and his Impossible Family came back to Earth to search for one of their missing children, they encountered Samuel who had since grown to find the life of a god boring and agreed to become part of the Impossible Family and took on the form of one of their children. He eventually returned with them to planet Poppup when they found their wayward child[199].

Meanwhile, the Lyja, the Skrull spy posing as Alicia Masters for months was uncovered thanks to the machinations of the Mad Thinker who deduced this situation and orchestrated a situation where Reed needed to do brain scans of those living in their headquarters the Four Freedoms Plaza[200]. When Lyja was revealed as a Skrull she revealed how she infiltrated the group but also admitted that she fell in love with the Human Torch in the process and was pregnant with his baby. The group then went out into space with Lyja to rescue the real Alicia from the Skrull empire. They clashed with Paibok who had used Skrull science to gain super-human abilities. Ultimately, the Fantastic Four managed to rescue the real Alicia. During the conflict Lyja seemingly sacrificed her life to save Johnny from danger and the Four left her body behind as Skull outpost exploded[137]. However, Pablok managed to escape in a life support vessel with the injured Lyja kept in stasis until he could get help[201].

Operation: Galactic Storm

Also around this time the Skrulls enemies the Kree and the Shi'ar were involved in a galactic conflict that was later dubbed Operation: Galactic Storm, a conflict that also drew in the Avengers as the war also threatened Earth[202]. Seeing this as an opportunity, the Skrulls began covert operations to escalate the conflict between the two sides. On the Shi'ar side, a Skrull posed as Empress Lilandra's advisor Araki, to encourage her to increase hostilities[203]. Another group of Skrulls then attacked a Shi'ar world in a Kree ship encuring the wraith of Gladiator who fought the Avengers, thinking them allied with the Kree. The Skrull pilots committed suicide before the Shi'ar could find out but Avengers Captain Marvel and Living Lightning discovered the truth before their ship was detonated[204]. The conflict escalated to the point where Lilandra sent a Nega-Bomb ship toward the Kree galaxy unaware that a Skrull flagship was heading to intercept it[205]. When the Kree Starforce teleported to the Shi'ar throneworld and attacked the Imperial Guard and Avengers, the Skrull posing as Araki attempted to assassinate Lilandra, but could not get the chance to do so. However the attack did prompt Lilandra to order the arming of the Nega-Bomb[206].

The Skrull posing as Araki then inflamed tensions between the Avengers and the Imperial Guard, and manipulating Lilanra into continuing hostilities toward the Kree. However a blast from the Living Lighting knocked "Araki" out, causing him to revert back to his Skrull form. Realizing that she had been manipulated into war by the Skrulls, Lilandra ordered a halt to the Nega-Bomb's transport between warpgates but by then it was already too late: the Skrulls had captured the Nega-Bomb with the Vision and Wonder Man trapped on board[203]. Passing through the Shi'ar warpgate, the Skrulls with the Nega-Bomb were impeded by Quasar, Her, and Binary but they were forced to allow the Skrulls advance when it was learned that the Nega-Bomb's proximity to Earth's sun was threatening to destroy it[207]. The Skrulls were opposed by Wonder Man and the Vision and while they succeeded in setting off the bomb, they were only able to do so in the outer fringes of the Kree galaxy. Although they failed in wiping out all the Kree, they slaughtered billions sending their age old enemies into utter chaos[208]}.

Renewed Aggressions

Eventually, Paibok's ship was rescued by Devos the Destroyer a being who sought galactic peace by destroying violent beings, even wiping out entire races. Paibok managed to convince Devos from trying to slay the Skrull empire by focusing on their mutual hatred of the Fantastic Four[201]. Soon Paibok and Devos became close allies, working to empower Lyja with the ability to shoot lasers from her fists while on their way to Earth[209]. They soon arrived on Earth just as Lyja was restored to full health[210].

Elsewhere in the universe another group of Skrulls were en route to wipe out the planet Dandesh in the Coalsack Nebula where the Xartans aka the Carbon-Copy Men, who had similar shape shifting powers due to the fact that the Skrulls found them inferior to themselves. The Xartans however tried to disguise themselves as the nearly extinct D'bari race to try and confuse their would be attackers[211]. Along their way to Dandesh they encounter the She-Hulk and her allies Weezie and US Archer on their quest to that same region to save their friends Razorback and Taryn O'Connell from the "D'bari". The Skrull commander agreed to help them due to the She-Hulk's slight resemblance to a Skrull[212]. Ultimately She-Hulk and her comrades rescued their friends and escaped while a D'bari petrification ray went off turning all the Xartans and Skrulls into stone[211].

Manwhile back on Earth, the Skrulls continued their plans for infiltration they began tracking the West Coast Avenger known as Mockingbird[213], and during the West Coast Avenger's battle with Ultron Mockingbird was replaced with a Skrull impostor who then infiltrated the group[214][215][216].[217] Mockingbird woke up on a Skrull world and found herself in a Skrull simulation of her life at the West Coast Avengers compound. But she soon saw through the ruse and broke free and escaped. She was pursued by a Skrull posing as her husband Hawkeye for years while she used her skills as a spy to survive[218]. While on the East Coast of Earth, having convinced Lyja that the Human Torch abandoned her and left her to die, the trio attacked Johnny at Empire State University. The shock of seeing his wife alive and well and a relentless assault forced Johnny to unleash a nova blast that forced his enemies to flee, but lit Empire State University on fire[219]. Wanted by the police, the Human Torch surrendered to the authorities, but Lyja and Paibok attacked the Torch at the court house, using their shapeshifting powers to make it seem the Torch had went berserk and forcing Johnny to flee from the law[220]. Johnny became a hunted fugitive and when he and the Fantastic Four clashed with the Secret Defenders, Lyja couldn't bring herself to hate her former husband any more and abandoned Paibok and Devos to help him[221].[222] Undaunted, Paibok and Devos next targeted Johnny Storm during his court trail over the fire at Empire State University, during the battle they joined forces with Klaw and Huntara who also sought to battle the Fantastic Four[223]. However the villains were thwarted by the Four with the help of their allies Spider-Man, Daredevil, Sandman and Silver Sable[224].

Also during this period, the Skrull who posed as the Contemplator resurfaced. Continuing his charade he joined Elders the Possessor, Obliterator, Judicator, Explorer, and Caregiver to investigate the strange embryonic space creature known as Origin. Their examination was interrupted by Quasar and his ally Thunderstrike, who had come to recover Quasar's missing Quantum Bands. The "Contemplator" grabbed one of the bands before Quasar could put it on, but Quasar managed to recover it by overloading the band blowing off the "Contemplator's" hand. Sharing his knowledge that he the "Contemplator" was an impostor, the Judicator used his powers to revert the impostor back to his Skrull form. Furious at this deception the Elders grabbed a Skrull and teleported away so that the impostor could face judgement[225]. His subsequent fate is unknown. The Skrull Empire was soon adversely effected by the so-called Infinity Crusade, when Adam Warlock's aspect of goodness, the Goddess was attempting to create universal peace. By this time the Skrull empire was lock in a battle with the Shi'ar, but the Goddess's influence caused both sides to cease fighting and depart[226]. Seeing what happened to his people and learning of the Goddess's involvement the Super-Skrull, seemingly unaffected, flew into space to hunt her down. Sensing the Super-Skrull's approach Goddess's assistant Moondragon dispatched Thor to battle the Super-Skrull, who was easily defeated[227]. However the Skrulls eventually returned to their hostile ways after the defeat of the Goddess at the hands of Adam Warlock, Thanos and Charles Xavier[228]. Shortly after this the Silver Surfer encountered a Skrull bounty hunter named Raze who was seeking revenge against his brother Vranz.[229][230]

Back on Earth, Paibok and his minions had regrouped[231], and attacked the Fantastic Four during a period in which they believed their leader Reed Richards was dead. Paibok and Devos captured the Invisible Woman, Thing, Human Torch and Lyja and brought them back to the Skrull homeworld where they were taken prisoner. However matters were complicated when Devos decided that was the time to lay waste to the Skrull empire and ordered his ship to destroy the Skull throneworld. During the chaos, the surviving members of the FF broke free and recovered a vial of Lacaroo, a medicine to help with Lyja's now complicated pregnancy and fled the Skrull homeworld. Meanwhile, branded a traitor by Sy'Bll, Paibok attempted to make amends by seemingly sacrificing his life to stop Devos's ship but hurtling it into sub-space[232]. Around this time, while on a mission with the West Coast Avengers, the Skrull posing as Mockingbird was killed battling Mephisto, Satannish and the Lethal Legion[233]. While in space, a Skrull known as Muraitak was a member of the Centurian Nova Corps. He was once ordered to bring Nova back to planet Xandar to battle the Lumphomoid known as Kraa.[234] After Nova helped defeaet Kraa, Muraitak allowed Nova to go free.[235] Elsewhere on Earth, thanks to the impulsiveness of the Impossible Man, invitations for the wedding of Rick Jones and Marlo Chandler in Las Vegas were issued to many of the Hulk and Rick's enemies including a delegation of the Kree and Skrulls. Despite some initial tension between the two delegations, Jones managed to convince them to sit on either sides of the wedding. Incidentally there was not a conflict between the two[236]. Among the Skrull party was Talos who drunkenly attempted to commit suicide to the cheers and adorations of his fellow Skrulls. The Hulk clashed with him, but realizing that Talos needed to restore his honor, the Hulk threw the fight making Talos a hero among his people again and restoring his will to live[171].

The Super-Skrull once again tried to attack the Silver Surfer not longer after this, against the orders of Empress S'Byll and was soundly defeated once more.[237] S'Byll has not been seen commanding the Skrulls since, her subsequent fate is unknown.

Infiltration of Earth

Following the fall of the Kree Empire, a group of Kree terrorists calling themselves Starstealth began raising havoc across the universe, including the planet Earth. A group of Starstealth terrorists were apprehended by the new group called Force Works and imprisoned in the Vault.[238] Part of the Skrull military became part of a war crimes tribunal against Starstrealth. Under the command of Major Hokk Algol they brokered an extradition tour with the Avengers. However their arrival coincided with members of Starstealth breaking out of prison with the assistance of Recorder RE-404.[239] The combined might of the Avengers, Force Works and the Skrull soldiers saw the escapees recaptured and extradited into space[240]. Other Skrulls active in the universe included Kerth a slave of the Broker,[241] and Morfex of the Cosmic Commandos.[242]

Around this time the Skrulls who had been turned into cows were sent off to slaughter by the United States government and the ground beef ended up in the food supply. Those who consumed the Skrull tainted beef became infected with a virus that was slowly killing them but endowed them with shape shifting powers even more advanced than the Skrulls themselves. Those infected were also capable of seeing Skrulls in their true forms no matter what shape they took. One person who consumed the tainted beef was apparently man known only as Ryder. His past is mostly a memory but Ryder has an unabashed hatred of Skrulls. Upon learning that others were infected by the tainted beef, he began going across the country killing Skrulls that he crossed paths with and recruited Skrull-meat infected individuals into his Skrull Kill Krew. Recruits included the English white supremacist known as Moonstomp, as California surfer Dice[243], New York punk rocker Riot[114], and super-model Catwalk[244]. The group discovered that Skrulls had infiltrated many places in society, and slew a Skrull who posted as Dice's girlfriend[243], Skrulls hiding within a New York night club and among the New York Police department[114]. Later the group got involved in freeing hostages from a plane hijacked by Hydra that contained Slovenian president Wassily Kurov. Ryder discovered Kurov was a Skrull but before he could slay him, he was stopped by Captain America[244]. After slaying Skrulls posing as the Fantastic Four[245] the Skrull Kill Krew discovered that the town of Pleasant Valley, Wisconsin was completely taken over by Skrull spies and wiped out the entire population of the town[246]. Floyd Donahue resurfaced shortly after this, offering to hire the Silver Surfer but was denied when the Surfer decided to travel back to Earth instead.[247] A short time later in deep space, a group of Skrulls attempted to obtain a millennias old comet that was returning to from the outer reaches of the universe, which was reported to have technology of monumental power. The Skrull fleet was wiped out by the bounty hunter White Raven who was seeking the comet for her employer Thanos. However her plans to obtain the comet were thwarted by the Silver Surfer.[248]. At some point Paibok resurfaced and allied himself with a number of Symbiotes battling both the Human Torch and Spider-Man[249].

On Earth a number of heroes, including the Fantastic Four and the Avengers were seemingly slain by the being known as Onslaught[250]. Following the seeming death of her husband, Lyja assumed a human identity and tried to forget about her past life[251]. However the heroes endured in a pocket universe on a world later dubbed Counter-Earth.[252]

Eventually the heroes were returned to their proper world and resumed their activities.[253] It was around this time that a Skrull named Sn'Tlo took control of a faction of Hydra, calling himself the Sensational Hydra. He then set about targeting Captain America by boosting his public image. To this end he orchestrated a number of victories for Cap such as a failed Hydra raid on the Smithsonian Institute,[254] and a battle with his old foe Batroc the Leaper.[255] Then when holding hostages at the top of the Empire State Building, Sn'Tlo and his minions captured Captain America at the heigh of his popularity and Sn'Tlo subdued him in Skrull bonds and then began posing as Captain America himself.[256] Sn'Tlo not only fooled the public, but also the Avengers with his ruse. Meanwhile, the real Captain America broke free from his bonds but was too late to stop his Skrull impostor from announcing to the country that one out of every twenty Americans is secretly a Skrull, causing massive riots as the American people gave into xenophobia.[257] Ultimately, Cap exposed the Skrull impostor and the civil unrest was soon ended.[258]

Not long after this a Skrull agent named Mrok was heading toward Earth on an undisclosed mission when he crossed paths with the Mithras a cosmic entity that brought peace to all living things it passed at the expense of free will and was reduced to a mindless state[259]. Ultimately the Mithras was defeated by the Earth mercenary known as Deadpool and all those effected by the Mithras were restored to normal[260]. Another Skrull agent disguised himself as Chicago's Officer Dragon in order to free Skrulls that were incarcerated on Earth. However this Skrull was exposed and apprehended by Nova.[261]

One group of Skrulls soon came under the command of Intelligencia a cybernetic collective mind that was the Skrulls' answer to the Kree Supreme Intelligence. Among the Skrulls under her command was the Super-Skrull. Intelligencia soon detected a threat from the human known as Kelly Kooliq who had become a walking nexus to all reality and could alter it to her whims. Under orders to kill Kooliq, the Super-Skrull finds himself unable to do so finding the task to be dishonourable leading to a clash with Captain Marvel, Moondragon and the Silver Surfer.[262] After battle between these heroes, the Super-Skrull managed to abscond with Kelly and return her to Skrell, the Skrull outpost where Intelligencia was located.[263] The three heroes tracked Kelly there and yet another battle broke out. Ultimately, Intelligencia decreed that so long as Kooliq continued to exist the universe would be at risk. Captain Marvel's alter ego Rick Jones then convinced Kooliq to end the threat by travelling to a reality wherein she never obtained her powers.[264] The Skrulls have not since been seen under the command of Intelligencia since, the subsequent fate of this construct is unrecorded.

The Twelve

Sometime later a group of Skrulls under the command of Granok came across the massive petrified form of the Living Monolith in space. Freeing the Monolith from his prison, the powerful mutate brought them to his leader Apocalypse who was at that time preparing to gather the Twelve a group of mutants that were fabled to shape the face of mutant kind. The Skrulls joined along with his plan over their mutual ideals on genetic purity and used this to further their studies on Skrull mutants.[265] It was about this time that the X-Men's years long trip from the destruction of the Skrull homeworld brought them back to Earth at their appropriate time. Their ship was jolted by electromagnetic burst that knocked the entire crew out.[121] They then replaced the X-Man Wolverine with a Skrull impostor who was virtually indistinguishable from the genuine article. The real Wolverine was then converted into Apocalypse's horseman Death.[266] For a time this impostor Wolverine worked along side the X-Men.[267] However suspecting a traitor in their ranks, Charles Xavier disbanded the X-Men in a elaborate plan to ferret out the impostor.[268] At the same time, two Skrulls posing as the deceased Mastermind and presumed dead Mesmero began trying to obtain the diaries of the late precognitive mutant Destiny which contained information about the gathering of the Twelve, they also spied on magnetic mutant Polaris to try and steal the jewel that adorned the original costume of her then presumed deceased boyfriend Havok. The two impostors failed to obtain the diary which was recovered by a team of X-Men who remained together.[269][270] Meanwhile, the Wolverine impostor was on a mission with another group of X-Men was ironically slain by Death, leaving the X-Men shattered by the belief that their long time comrade was killed.[271] Knowing that at least a single impostor was exposed, Xavier called back his X-Men and with Phoenix administered a psychic test to determine that none of the other members of the team had been replaced, they then revealed that the Wolverine who fought and died beside them was really a Skrull impostor. During this revelation another another Skrull attempted to replace Archangel but that attempt was also thwarted[272].

In Egypt, Garnok and his minions continued to experiment to make an even perfect Wolverine impostor to little effect, while Fiz one of the mutant Skrulls attempted to get his fellow outcasts to rebel against their masters, inspired by the legends of how Xavier once tried to save their homeworld. The Skrulls then attempted to kidnap Polaris and recover the gem to Havok's costume. They succeed in capturing the gem, but thanks to a trick involving an image inducer, they really brought the X-Men's field leader Cyclops into their headquarters, luring the reset of the X-Men to their location. During the course of the battle, Death appeared and slew Garnok and the X-Men learned his true identity before the Skrulls escaped.[125] The mutant Skrull Fiz managed to break out of his cell during the attack and followed the X-Men back to their base. There he appealed to them to help free his fellow mutants offering to bring them right to Apocalypse's base[265]. When a squad of X-Men, including the uncaptured members of the Twelve arrived outside Apocalypse's base, the Skrulls, Apocalypse's minions and his Four Horsemen defended the base. Skrulls then posing as Pestilence, Famine, Astra, Joseph, Havok, Colossus, and Cannonball managed to capture Storm, Magneto, Bishop, Polaris, Cyclops, Phoenix, and Professor X, the last members of the Twelve that required capturing[273]. The Skrulls kept the remaining X-Men at bay long enough for Apocalypse to drain the required energies to transfer his essence into a new body. While he intended to take the body of the X-Man, Cyclops jumped in the way and Apocalypse took over his body instead[274]

With his newfound power Apocalypse tried to replace the Earth-616 reality with those of Earth-23378,[275] Earth-2203,[276] Earth-2841,[277] Earth-32000[278] and Earth-32098. Many of these realities also included the Skrulls and Apocalypse until ultimately the X-Men succeeded and restored reality back to normal. However Apocalypse, and presumably his Skrull allies, escaped.[279] With the Skrull mutants needed guidance and mentorship, Charles Xavier agreed to return to space with them to train them to be freedom fighters for their own people, this group of mutant Skrull champions became known as Cadre K named after the Skrull classification for their breeding class.[280]

Maximum Security

The Skrulls empire soon became involved in the manipulation of the Supreme Intelligence once again. After obtaining the Forever Crystal[281] he used it to hyper-evolve a number of the Kree into an evolved state.[282][283] Calling themselves the Ruul, these evolved Kree then posed as a new race offering advanced technology to other alien races to gain their trust, including the Skrulls. About this time Cadre K had become notorious among the Skrulls for liberating other mutant Skrulls and fighting against the empire. This prompted Ambassador Kreddick the Skrull representative of the Intergalactic Council to demand something be done about human interference in the affairs of the universe. After an attack by Ego the Living Planet was thwarted by Cadre K and the Silver Surfer, manipulations by the Ruul and insistance from the Skrull ambassador convinced the Intergalactic Council to turn the planet Earth into a prison planet, dumping intergalactic criminals there as well as preventing the people from Earth from leaving and influencing the universe[284].

This put the heroes of Earth on the defensive while their planet was invaded by armies of alien criminals[285]. Cadre K, led by Xavier soon rescued Bishop from the clutches of Deathbird and sent him on to Earth to recover their missing member Z'cann[286]. On Earth, Z'Cann attempted to locate the X-Men and was injured by the Blood Brothers. In an attempt to tell the group what was going on she made physical contact with Rogue allowing her to absorb her memories and powers, altering Rogue's powers.[287] Meanwhile, Xavier and Cadre K decided it was time for more overt operations[288] and attacked the Intergalactic Council directly but were captured as a result.[289] Ultimately, Cadre K were instrumental in exposing the Supreme Intelligent's plot and Earth's heroes eventually triumphed, forcing the Intergalactic Council to lift their restrictions off the Earth. In the aftermath, Xavier had decided that Cadre K had proven themselves and no longer needed his guidance and sent them off to fight for their people[283].

Laying Low

What followed was a period of what appeared to be a low ebb in Skrull activity for a time as the Skrulls appeared to be involved in less grandiose schemes or plans of conquest. A group of Skrulls were assigned to assassinate the Super-Skrull who was hiding out on Earth. However this plan was thwarted by the Human Torch.[290] A pair of Skrulls piloted Thanos to investigate a strange black hole that appeared and disappeared, and were slain as part of a complex plot by the Thanosi to slay their creator, a plan that failed.[291]

Paibok returned once again, seeking to boost his powers using Centauri technology, however the process left him disfigured in his normal Skrull form. He then gathered a band of Skrull outcasts to obtain Talmadge a young Watcher that was being held by the Kree in the town of Clarkston located in the State of Michigan[292]. To this end Paibok and his minions disguised themselves as members of a freakshow that operated in the town. However their plans were complicated thanks to the involvement of the Thing who happened to come to the town[293][294][292]. After a pitched battle, the Skrulls were defeated and Paibok was taken prisoner by the Kree[295]. Also during this time frame, a group of Skrulls lived peacefully along side a group of Krees who were living in secret in the town of Raven's Perch, New Jersey home of the hero group known as the B-Sides[296][297][298].

Around this time the Grandmaster entered into a competition with Krona a cosmic being from a distant cosmos to pit their realities greatest champions against one another. In order trick both the Avengers and the Justice League of America into learning of each others worlds, both Grandmaster and Krona made it appear that both realities were being invaded by beings from the opposing universe. In one particular instance a Skrull invasion fleet was teleported to the planet Thanagar in that other world where the clashed with the Thanagarian Hawkmen[299]. Ultimately these Skrulls were returned to their native reality before the contest and resulting crisis that threatened both realities was thwarted by the Avengers and the JLA.[300]

While not involving the Skrulls directly, a number of events occurred on Earth that would lay the path to a planned invasion of Earth: the dissolution of the Avengers after a devastating battle against the Scarlet Witch who had temporarily gone mad,[301] and the apparent death of the Asgardian thunder god Thor and destruction of Asgard.[302]

Not long after this another Skrull who was sent to Earth to infiltrate the super-hero community was Z'Reg to infiltrate the Avengers ranks.[303][304] However he arrived shortly after the Avengers had recently disbanded.[301] Z'Reg soon came to love Earth and abandoned his mission taking on a human guise and living as the unassuming next door neighbour of Curtis Doyle.[42] Also following the dissolution of the Avengers the time traveller known as Iron Lad came to the past to recruit a new team called the Young Avengers, among the team included Teddy Altman—who unknown even to himself at that time was really Dorrek VIII, the long lost heir to the Skrull throne[305]. Teddy served with the Young Avengers for a time as Hulkling, unaware of his true heritage.[306]

Another event playing into the events that followed was the revelations that Nick Fury engaged in an illegal act of war against the nation of Latveria using many of America's greatest heroes. This violation of international law forced Nick Fury to go underground to avoid capture.[307].

Pre-Invasion Phase 1

By this time the Skrulls soon sought out Veranke having come to believe in the prophecy that the Skrull empire were was to inherit the Earth. Varanke was then returned to Satraini and named the queen of the empire. She was brought before Skrull scientist Dro'ge who had perfected the technology to engineer Skrulls to not only duplicate appearances, but to duplicate them down to a cellar level. In addition to this Skrulls were given the power to duplicate the super-human abilities of those they are replicating. Through specially mental conditioning they would be programmed with the memories of their target and not only have their memories but believe that they were that person. The process was so thorough those under deep cover would be undetectable through technological, telepathic or mystic means. He had initially sent a Skrull named Siri to pose as the assassin Elektra[75]. With the real Elektra in their captivity, they experimented on her to try and learn the secret of her many resurrections, but to no avail[308]. She was soon brought back to Satraini and brought before Veranke[309]. So impressed by this, Veranke asked to be one of the first deep cover agents and chose to replace the Earth heroine known as Spider-Woman[75]. Before leaving for Earth, Veranke paid a visit to planet Tarnax VII and ordered her loyal follower Chrell to eliminate Dorrek VIII when they launch their invasion so as not to have his opposition with her invasion plans[310]. At this point, Skrulls had once more infiltrated the terrorist organization known as Hydra. Using one of their agents they offered Spider-Woman the chance of having her powers restored in exchange for being a double agent for Hydra[311]. She agreed to the offer, under the direction of SHIELD director Nick Fury so they could expose and shut down that Hydra cell[312]. However, instead she was replaced with Veranke who promptly took Spider-Woman's place[313]. It was also around this time that the Skrulls managed to track down Lyja who had been living in her human guise for years. They convinced her to join their cause to get revenge against the Human Torch for abandoning her after he returned after his "death" at the hands of Onslaught, and used Skrull science to restore her laser-fist powers.[251]

Veranke then began meeting with other Skrull infiltrators that were slowly infiltrating both Hydra and SHIELD. They then began planning three major plans: The first was to eliminate the rare metal known as Vibranium as the weapons made from it would be unbeatable. Veranke ordered a team of SHIELD infiltrators to mine the supply found in the Savage Land region of Antartica and making it appear that it was being carried out by Nick Fury loyalists. Another concern was the growing mutant population, prompting the Skrulls to prompt a campaign to fan the fear of anti-mutant bigotry in the hopes it would decimate the mutant population. Lastly they also began focusing their attention to more powerful humans such as the Scarlet Witch, Doctor Doom, Hercules and members of the Illuminati. However Veranke put orders that Reed Richards and Tony Stark would not be killed until after their invasion was complete and they could be put on trail and executed on the Skrull homeworld[313]. Shortly after, Veranke had many other Skrulls go through the process to pose as various heroes and villains from Earth, with false memories leaving them to believe that they were kidnapped by Skrulls at various points during Earth's most recent history and were placed aboard a prisoner ship where it was planned they would escape once the planned invasion was ready to begin[314]. Shortly after this, the Skrulls located Nick Fury hiding out in Mexico. They sent a Skrull posing as Valentina Allegra de Fontaine, however Fury soon uncovered that she was a Skrull and slew her. Fury then became aware of the Skrull invasion and went deeper under ground to prepare to stop it[315]. The Skrulls next went after the assassin Elektra, capturing her and having her replaced by a Skrull named Pagon. Posing as Elektra, Pagon then slew the leader of the assassins known as the Hand, taking leadership of them securing yet another power for the Skrulls[316].

Through her connections within SHIELD, Veranke continued to pose as Spider-Woman, eventually taking part in stopping an attempted break out at the Raft prison in New York where she assisted Captain America, Iron Man, Luke Cage, Spider-Man, and Wolverine in stopping the escape of many of the super-powered inmates there[317][318]. This breakout was facilitated by the criminal known as Electro, who was secretly paid by the Skrull posing as Elektra to do so, so that the Skrulls could secure the use of mutate Karl Lykos to use to find the Vabranium hidden in the Savage Land[316]. When Iron Man and Captain America formed the New Avengers, "Spider-Woman" was selected as a member and from this position she would continue to manipulate Earth's mightiest heroes until it was time for the Skrull invasion of Earth[319]. Ironically enough, the New Avengers mission was to recapture the criminals who escaped from the Raft, starting with Karl Lykos[320]. About this time the Skrulls posing as SHIELD agents were discovered by the Savage Land's ruler Ka-Zar and his wife Shanna who uncovered who they really wear. Overhearing that the Skrulls were busy infiltrating the United States government, SHIELD, and the Avengers, when the New Avengers arrived they did not approach as they did not know whom they could trust[321]. The New Avengers eventually uncovered the illegal SHIELD operation there[313], however the Skrulls used a SHIELD helicarrier to destroy all evidence before the Avengers could learn of Skrull involvement, and removing the Antarctic Vabranium as a potential weapon against their invasion[322][313]. Meanwhile Veranke continued to fight alongside the Avengers leaving them unaware of her Skrull nature.[323]

After years away, Titannus resurfaced having been brainwashed into being an agent for the Trellians. Left to believe he was escaping an oppressive ruler with Princess Amissa as his unwilling paramour to Earth,[324] where be began monitoring Earth's heroes.[325] His presence was soon discovered and he Titannus clashed with the Japanese mutant Sunfire[326][327] a battle that left Tokyo decimated.[328][329] This ultimately led to a clash between Titannus, Spider-Man, Wolverine, Dr. Strange, Nova, Hulk, She-Hulk and Ms. Marvel.[330] The heroes quickly freed Amissa who revealed that she was not Titannus' lover and that he kidnapped and that her father was not a tyrant. Distraught and mentally unstable, Titannus blew his own head off, ending the battle. However his enhanced healing caused his head to rapidly regenerate.[331]

Not long after this the Super-Skrull learned of Dorrek VIII's true heritage and came after the boy, kidnapping him from his fellow Young Avengers after revealing to Hulkling his Skrull heritage, seeking him to take his place as ruler of the Skrull empire[332]. However Super-Skrull was not only tracked down by the Young Avengers, but also the Kree who sought to take the boy to rule their empire as well[73]. This resulted in a battle between the Young Avengers, New Avengers, and the Kree and Skrull entourages coming to claim Hulkling[72]. Ultimately, Teddy and the Super-Skrull came up with a ruse to stop the fighting: The Super-Skrull, posing as Hulking agreed to leave to spend half his time ruling the Kree and the other half the Skrulls left with both alien envoys allowing Hulkling to stay on Earth and resume his activities with the Young Avengers.[333]

Z'Reg resurfaced eventually, after his neighbour discovered a powerful ring fashioned out of shards of the Cosmic Cube he became the costumed hero Freedom Ring, but was crippled in a battle.[42][304] Z'Reg revealed himself and decided to teach Curtis how to utilize the ring. Taking on the costumed identity of the Crusader, Z'Reg also helped Freedom Ring train.[303] Eventually the duo assisted the New Avengers in battling the Iron Maniac. Freedom Ring died stopping the Iron Maniac, and the Crusader took his ring and continued his costumed heroics.[334] Titannus escaped shortly after this and went on a rampage again, Crusader joined a number of heroes in stopping him and he was eventually detained and taken into S.H.I.E.L.D. custody.[335] By this time Ethan Edwards, now an adult working for the Daily Bugle newspaper in New York began using his powers as a costumed hero called Virtue. He joined Spider-Man in fighting the Absorbing Man,[336] the two learned of their alter egos and Spider-Man agreed to help Virtue train to be a better hero and began learning about his origins[127]. Virtue eventually brought the Fantastic Four back to his "hometown" of Hicksville where they found the ship that brought him to Earth and learned of his Skrull heritage[126]. The ship then began encoding Virtue with his mission perimeters to take over the Earth leading to a clash with the FF, the conflicting memories of his Skrull mission and his Earthly upbringing causing a massive and violent identity crisis[337]. The New Avengers managed to detain Virtue and Spider-Man convinced him that despite his true heritage he was still a good person. Virtue then decided to abandon costume heroics and use his healing powers, which came from some unknown source outside of his fathers genetic tampering, to heal the sick and the infirm[338].

Pre-Invasion Phase 2

During this period the Skrulls were locked in a conflict with the Majesdanians. An order of Super-Skrulls were then trained, which included a young Skrullian prince named Xavin.[339] He was trained under the command of a Skrull trained named Chrell[340][341]. In order to try and broker peace an arranged marriage was agreed upon by where Majesdian parents Frank and Leslie Dean agreed that their daughter Karolina would marry Skrull prince Xavin. Xavin eventually came seeking Karolina[339] leading to a battle between Xavin and her teammates the Runaways. Ultimately, Karolina agreed to go into space to marry Xavin in order to broker peace between the two races.[342] However this plan did not work and eventually Xavin and Karolina returned to Earth and reunited with the Runaways.[343][344] A prison ship carrying Paibok the Power Skrull and other alien criminals crashed in the town of Hoot's Bluff, Alaska.[345] There the escapees clashed with Drax the Destroyer until they were apprehended again.[346][347][348].

Veranke soon got the opportunity to deal with the Scarlet Witch, whom the Avengers were grouping to deal with after her mental instability continued to cause her to warp reality. When the Avengers called in veteran members such as Yellowjacket, they were still unaware that both Spider-Woman and Yellowjacket were Skrull impostors. Veranke and Criti Noll plotted to eliminate the Scarlet Witch once and for all once they arrived in Genosha where she was staying in convalescence[313]. However they did not get the chance as the encounter led to a battle between the New Avengers and X-Men against the Scarlet Witch's family: her father Magneto and brother Quicksilver. Ultimately, the Witched used her powers to replace Earth-616 with Earth-58163[349]. Because they were Skrull impostors, Veranke and Criti Noll were one of the few beings in this new reality that retained their memories of the old one and plotted to do away with once and for all[350]. However in the final battle, the Scarlet Witch realized that mutants were the cause of constant conflict and restored reality back to normal, however she removed the mutant gene from 99% of the mutant population[351].

Also around this time a number of Warskrulls were still active at this time. One named Flaw fought alongside the Shi'ar Death Commandos who were assigned to eliminate the entire Grey family genome.[352].[353] The X-Men also clashed with Warskrulls who were trapped on a space station crippled by the powerful mutant Vulcan[354]. Another Skrull was stopped from using nuclear arms on an Earth space station by Doc Samson.[355] This Skrull believed himself to be one of the Skrulls who Reed Richards turned into a cow years earlier, but this claim has not been substantiated and contradicts previous reports that all the Skrull Cows were ground up into hamburger.


When it was learned that the positive matter universe was expanding and consuming the so-called Negative Zone, it's ruler Annihilus launched a massive full scale assault on the positive matter universe that was commonly referred to as the Annihilation Wave. It burst through from the centre of the universe referred to as The Crunch and began decimating everything in their path. They destroyed the Klyn prison located at the centre of the universe near what is called "The Crunch". Among the many prisoners Piabok was a prisoner there. The planet Xandar and the Nova Corps were decimated in the opening salvo of what would be called the Annihilation War[356]. Paibok and the other prisoners from the Klyn that were found useful were made slaves of the Annihilation Wave.[357]

On the 15th day of the Annihilation Wave reached the Skrull galaxy and began laying waste to it. This destruction was felt all the way on Earth by Veranke and her followers who had met to discuss the after effects of decimation of the mutant race. Feeling the genocide of billions of her people convinced Veranke that yet another prophecy from the Book of Worlds was coming true and that they should stay on their course[350]. Within 17 days the Annihilation Wave pushed through the Andromeda Galaxy cutting a swath through the Skrull Empire. One of the first recorded decimations occurred at Tala Prime, leaving no Skrull survivors[358]. By day 19 the Wave cut through Tercera Prime and began to swarm the Eudorian Reach[359]. On day 20 they pressed into Tecera Minor in the Peripheral System the centre of a battle between the Annihilation Wave, Galactus and his Heralds[360], a battle that ended in the Badoon/Skrull Neutral Zone known as the Obad'an Rim[361].

On day 41 of the Annihilation Wave, the forces of Annihilus reached the Skrullian world of Ask'lo. Despite the addition of the Super-Skrull to the Skrull armies there they proved no match for the Annihilation Wave's most powerful weapon, the planet destroying Harvester of Sorrow which destroyed the planet. Escaping alive, and on day 46, Kl'rt attempted to warn Baroness S'Bak and demanded that she allow him to lead an attack on the Harvester of Sorrows. This was because it was headed to the Skrull world of Zaragz'na the world where Kl'rt's son lived. The Baroness refused the Super-Skrull and insulted her, angering him enough to strike her. She ordered her minions to slay him, but the Super-Skrull managed to escaped thanks to R'Kin a young Skrull mechanic whose father fought along side the Super-Skrull years earlier. Seeking to find a weakness in the Harvester, Kl'rt and R'Kin travelled to Earth and convinced Reed Richards to allow them to travel into the Negative Zone through his portal there[362]. For the next seven days they travelled through the Zone, Super-Skrull slaying all those that did not provide him with answers, disturbing R'Kin. On day 56 in the Nexus of Igoa they found UX-73 a Negative Zone prison to find Hawal the scientist who invented the Harvester of Sorrows. He captured Hawal and gathered an army of prisoners as his army, including the mechanoid known as Praxagora and Preak[363]. When the Super-Skrull led his warriors back into the positive matter universe to stop the Harvester of Sorrows from destroying Zaragz'na, they were suddenly betrayed by R'Kin, who had been twisted by the Super-Skrull's brutal ways that he decided to side with the Annihilation Wave and helped them take Super-Skrull and his allies prisoner and Zaragz'na was destroyed[364]. Kl'rt broke free and in the ensuing battle, the Harvester was destroyed and Kl'rt seemingly slain in the explosion[365]. By day 79, the Skrull Empire was no more after 64 days of non-stop invasion[366].

Other Skrulls active during the Annihilation War was Talos who now lived in the fringe Kree world of Godthab Omega. On the 95th day of the Annihilation War he clashed with Devos, who had tracked Talos in order to slay him[367]. The pair were captured by Galorian the former disciple of the Shaper of Worlds who sought to remake Godthab Omega into a paradise[368]. The planet was soon devastated by the Annihilation Wave on the 99th day of the Wave and in the aftermath Ronan the Accuser ordered both Talos and Devos to work together to help the survivors get off planet[369]. Devos was later seen fighting along side the the United Front, warriors from various empires in the galaxy under the command of Nova who were working together to stop Annihilus and his forces. He was involved in the fighting on Daedalus 5 in the Kree galaxy[366].

On day 212 of the Annihilation War, Praxagora joined the United Front, bringing with her the seemingly deceased body of the Super-Skrull when they were suddenly attacked by Ravenous and his army of Centurions, an elite cadre of super-powered warriors under Annihilus' command[357]. Kl'rt was revived by the United Front's healing machines and assisted in the evacuation of Deadalus 5 as it fell under the Annihilation Wave[370]. When Ronan the Accuser decided to rest control of the Kree Empire from the House of Fiyero, the Super-Skrull and Praxagora convinced him to allow them to accompany him[371]. On the Kree world of Hala, Super-Skrull assisted Ronan in exposing the House of Fiyero as working with the Annihilation Wave and helped him retake his homeworld[372]. Following the eventual slaying of Annihilus and the defeat of his Annihilation Wave, officials believed that the Skrull Empire was destroyed and showed no sign of recover[373]. Super-Skrull continued to work alongside Ronan for a time[374].[375] Freed from slavery under the Annihlation Wave, Paibok briefly accompanied his fellow slaves the Delinquent and Terrax to New Xeron.[376]

Pre-Invasion Phase 3

While the Annihilation Wave was on going, on Earth, Veranke and her followers continued on their mission to invade the Earth, putting their final plans into motion. Meanwhile on Earth, with Skrull forces slowly infiltrating all levels of American government and various terrorist organizations, Veranke and her faithful follows slowly sewed the seeds of dissent among the ranks of Earth's super-heroes.

They then began eliminating the last beings that would stand in their way. The Skrull posing as Jarvis used his position to access files on the mental health of the Sentry, whose sanity was questionable at the time. The Skrulls then began manipulating his therapy sessions so they could use his mental illness against him when he began to get into the way of their plans[102]. The Skrulls were either directly, or indirectly involved in the exile of the Hulk from Earth[377] as well as various other super-human related disasters the Superhuman Registration Act was passed in the United States forcing superhumans and costumed crime fighters to register with the government and reveal their identities. This split the super-hero community in half and two opposing sides battled in a conflict that was referred to as the Civil War[378]. In the middle of the war, a Skrull posing as the deceased Captain Marvel[379] appeared in the middle of the conflict. Leading the heroes to believe he was plucked out of the past and was still dying of cancer, "Mar-Vell" agreed to remain in the present and help run security Prison 42, a prison constructed in the Negative Zone to hold those who refused to register with the government[380]. The conflict ended with the surrender of Captain America, the leader of the anti-registration resisters[381]. Captain America was later seeming assassinated while in custody by one of his foes.[382] On the night Captain America was murdered, SHIELD operatives Dum Dum Dugan and were replaced by the Skrull that was posing as the Countess[383]. Criti Noll however soon became affected by the Henry Pym implant in his mind and tried to warn an Oregon restaurant owner of the coming invasion. This was stopped by the Dugan impersonator who slew Citri and had his forces eliminate all eye witnesses of the battle. Citri was then replaced with a clone that had none of Pym's capacity for human compassion[384].

While in Britain at that time, John the Skrull resurfaced once again, having been working for years under the British government's newest paranormal investigation agency MI13 working with the mutant Pete Wisdom in defending the British Isles from various threats[385].[386] Not long after this John reunited the Skrull Beatles to assist in stopping the Martian Masters of Earth-691 from invading Earth-616 through a portal opened in London.[387]

Following the end of the Civil War, Iron Man was appointed the new director of SHIELD and he, "Yellowjacket" and Reed Richards began the 50-State Initiative[388] a government program that trained super-humans to use their powers properly and place them in government sanctioned super-hero groups in every state in America. Meanwhile, Iron Man also formed the so-called Mighty Avengers, a government sanctioned version of the group[389]. Veranke initially remained an unregistered super-hero in her guise of Spider-Woman fighting along side the outlaw New Avengers. Meanwhile the new Criti Noll used her position as "Yellowjacket" to place Skrull agents with in the Initiative, while Iron Man's use of StarkTech in all measures of SHIELD defences gave the Skrull infiltrators access to the entire defence of the United States.[388][390]

Another Skrull active on Earth, but not involved in the planned invasion was Jazinda Kl'rt Spawn, the daughter of Kl'rt the Super-Skrull. Who unspecified years earlier attempted to steal the Sy-Torak gem from the Kree empire for the Skrulls. During a skirmish she attempted to swallow the gem which endowed her with incredible regenerative powers, but it was permanently bonded to her body. Refusing to sacrifice her life to have the gem extracted Jazinda fled the empire and her father vowed to track her down and kill her.[391] Fleeing to Earth Jazinda became a bounty hunter in America,[392] eventually working along side the She-Hulk.[393]

Also around this time the Skrulls on Kral IV had finally rebuilt their society. Under the leadership of a Skrull calling himself Lou "Biggie" Smalls things mostly went back to the gangster themed status quo. However some of the Skrulls were repentant of their former slaving ways and inspired by television transmissions of the American Civil Rights movement of the 1950s and 60s, adopted the forms of African-Americans. Their leaders took on the guises of Martin Luther King, and Malcom X and attempted to change the way of their people through peaceful demonstration. Biggie Small and his minions fought against this and the African-American Skrulls were put on the defensive in the area of Kral fashioned to resemble New York City's Harlem district. Finding that passive resistance was not working Malcom X then formed the Black Panther Party milita to fight against the Skrull slavemasters.[394] More recently the Fantastic Four (now counting the Black Panther and Storm among the roster) were transported to Kral by King Solomon's Frogs where they were instantly attacked by Biggie Small's minions.[395] The Skrull Slavers managed to capture the Thing, Human Torch and Black Panther while Storm managed to escape into "Harlem" and join up with the Black Panther Party. The three captives were then forced to fight in the arenas against alien creatures and each other.[396] Although the Black Panther Party resistance was almost betrayed by the double crossing "Bumpy" Johnson, Storm managed to free her comrades and Biggie's regime was demolished allowing Martin and his followers to rebuild a more peaceful society. The Fantastic Four were later teleported back home by King Solomon's Frogs.[394]

Following the Inhuman's so-called Silent War with the Earth over stolen Terrigen Crystals[397], the Inhuman leader Black Bolt was captured and replace with a Skrull impostor[398]. Soon the Hulk returned from his exile after a harrowing experience on the planet Skaar where he became their warrior king only for the warp core of the ship that brought him there destroyed the planet.[399] Seeking to get revenge against the Illuminati, the Hulk and his Warbound first landed on the moon where he brutally beat "Black Bolt" before turning his rage toward the Earth.[400] Black Bolt's queen Medusa nursed the impostor back to health, further entrenching his infiltration into Inhuman society.[398] The Hulk meanwhile was soon taken down by Earth's heroes, although at the cost of Dr. Strange's hands being crippled—thus limiting his magical powers.[401] In the aftermath of the battle, Skrulls had Amadeus Cho's pet coyote Kurby replaced with a Skrull as well.[8] This conflict also removed two more players that would work against the Skrulls planned invasion. Skrulls also attempted to infiltrate the Hood's super-criminal gang by trying to replace Madame Masque. However the Skrulls posing as SHIELD agents who attempted to cause this infiltration failed to do so and when the Hood uncovered this secret plot, and exposing a Skrull posing as the Slug. Exposing this infiltration helped cement loyalty from the Hood's minions.[402]

Secret Invasion: Infiltration

Later, the New Avengers clashed with the Hand in Japan. At that time the organization was still under the command of Pagon while posting as Elektra. She clashed with the outlaw New Avengers[403][404][309][405]. Ultimately the battle ended when the New Avenger known as Echo slew Pagon, revealing her Skrull identity[406]. While taking the corpse of Pagon back to the United States. When "Spider-Woman" suggested they bring the corpse to Tony Stark, the rest of the team refused. However, an opertune plane crash allowed Spider-Woman to escape with the corpse[407]. Veranke then delivered the corpse of Pagon to Iron Man and his Mighty Avengers, with news of how her former comrades believed that there was an impending Skrull invasion, cementing her "loyalty" to the Mighty Avengers[408]. Iron Man brought the corpse of Pagon to his comrades in the Illuminati to puzzle over a completely undetectable Skrull in their midsts and the implications of a wider invasion. It was during this meeting that the Skrull posing as Black Bolt then attacked with another Super-Skrull warrior. The heroes just barely managed to to stop these Skrull attackers slaying them both, however the Illuminati disbanded unable to trust themselves let alone anyone else.[409] Iron Man turned over the corpse of the Skrull who posed as Black Bolt over to his wife Medusa, who was furious to learn this.[410]

Also at this time, "Yellowjacket's" work in the Initiative saw the graduation of a number of recruits and the establishment of super-hero teams in various states in America, each group that was formed by "Yellowjacket" included at least one Skrull impostor. In Arizona the Desert Stars were infiltrated by the Blacksmith[411]; In Arkansas, the Battalion was infiltrated by a Buford Hollis (Skrull) (Earth-616)Skrull posing as Razorback[412]; Florida's Command was planted with a Devil Dinosaur impostor[412]; The Calvary of Georgia was infiltrated by a Skrull posing as Thor Girl[413]; Hawaii's Point Men had a Skrull posing as a novice hero named Magnitude[390]; a Frog Man impostor was among the ranks of Kentucy's Action Pack[414]; In addition to a Skrull posing as Equinox in Montana's Freedom Force[413]; New Mexico's Mavericks team had a Skrull posing as She-Thing[411]; An unidentified Skrull infiltrated Nevada's Heavy Hitters team[415]; While Pennsylvania's Liberteens were infiltrated by the Revolutionary[416]; Oddly a Skrull chose to pose as Red Wolf's pet Lobo among the Rangers of Texas[412]; A Skrull posed as Skyhawk of Washington State's Earth Force[412]; and lastly the Grasshopper was a Skrull spy within the Wisconsin's Great Lakes Initiative team.[412]. Ironically, one Initiative recruit, the Crusader[417], while a Skrull, was not part of the invasion plans and completely unaware of them, but soon began to suspect that "Yellowjacket" was a Skrull agent, making him to question his loyalties, while also fearing being exposed himself.[390]

By this time the Skrull posing as Captain Marvel had developed a religion based on his adopted crusade for peace. On a mission with Ms. Marvel they uncovered Skrulls posing as Kree soldiers unnerving "Mar-Vell".[418] This so unnerved Mar-Vell he questioned a Skrull who captured while impersonating the deceased villain known as Cobalt Man, who revealed to "Mar-Vell" of his involvement in the coming Skrull invasion of Earth;[419] After a brief personality crisis, "Captain Marvel" decided to accept his adopted Kree heritage and continue being the hero he was modelled to follow after.[379] Also during this period an attempt by Super-Skrulls to take down and replace Ms. Marvel also failed.[60][420][421] In addition to this, the Skrulls captured Beta Ray Bill to utilize his mystical hammer Stormbreaker as a weapon against the Asgardian gods.[422]

Nogor soon arrived on Earth and seeing the mutant Darwin as the missing link between mutants and Skrulls posed as the mutant known as Longshot and attempted to lull Darwin into a sense of false security.[423] Nogor's presence on Earth was detected by Jazinda who went after Nogor in Chicago with She-Hulk, which led to a brief conflict with X-Factor Investigations who were searching for Darwin.[9] Eventually the heroes put aside their differences and were able to capture Nogor and with the help of Darwin knock him out. Jazinda and She-Hulk then took Nogor as their prisoner in the hopes that they could use him as leverage to stop the coming invasion of Earth.[424]

Secret Invasion

Finally, Veranke and her followers in the Dard'van launched their jihad against the planet Earth. While Iron Man presented the corpse of Pagon to Reed Richards and "Henry Pym", the Skrull posing as Dum Dum Dugan paid a visit to the S.W.O.R.D. space station just as a Skrull ship was on a crash course with the Savage Land. With the Mighty Avengers heading to the Savage Land, "Spider-Woman" tipped off the New Avengers as well to get both groups into a conflict there. "Dum Dum" and his minions then detonated the SWORD space station, while "Jarvis" activated a virus that disabled all Starktech on the planet, crippling not only Iron Man, S.H.I.E.L.D., prisons like the Raft and the Cube, and the majority of Earth's defences and even first response emergency rescue. The Skrull posing a Yellow Jacket then revealed herself as well, taking Reed Richards prisoner.[425]

What followed was a simultaneous series of attacks in what is now referred to as the Secret Invasion of Earth. As all attacks happened simultaneously, and for the sake of clarity each attack has been detailed separately below based on the geographic locations.

Savage Land

In the Savage Land, the New Avengers and Mighty Avengers clashed until the Skrull virus began to infect Iron Man's armor, stopping the fight[425]. Then the crashed Skrull ship opened revealing a number of Super-Skrulls in the forms of various heroes, mentally conditioned to believe that they were the real heroes and were captured at various points in the past[314]. Unable to tell who Skrull impostor from the real heroes, a fight soon broke out between the Avengers and the "heroes" who emerged from the ship. During the battle, a Skrull posing as the Vision took advantage of the Skrulls manipulations of the Sentry's therapy[102], convincing the Sentry that the battle was being caused by his evil persona the Void. This caused the mentally fractured hero to flee the battle. Iron Man had Ms. Marvel pull him out of the battle so he could try to repair his armor, unaware that Veranke was following them. Meanwhile a sudden attack by the Tyrannosaur sent hero and Skrull impostor scattered. One Skrull impostor was posing as Mockingbird managed to convince Ronin that she was his seemingly deceased wife[426]. Spider-Man ran into Ka-Zar and Shanna and learned about the Skrull's involvement in the New Avenger's earliest mission in the Savage Land when they were suddenly attacked by a Skrull posing as Captain America[321]. While elsewhere Wonder Man clashed with a Skrull posing as his long time friend the Beast.[427]

Having sent Ms. Marvel to New York to get help, Iron Man was then confronted by "Spider-Woman" who then tried to convince Iron Man that he was really a Skrull infiltrator who helped with the invasion effort[428]. The Black Widow slew the Skrulls posing as the Beast and Phoenix and came to Iron Man's aid, sending Veranke fleeing[429]. However with no means of telling the real heroes from the impostors and with Iron Man fighting off the effects of the Skrull virus the situation was struck in a tense stalemate[430].

Thunderbolt Mountain

At the start of the Invasion, Khn'nr the Skrull posing as Captain Marvel was still conflicted with his false Kree memories and his orders in the Skrull invasion. Finally giving into his mission orders[427]. Following these orders he launched an attack on the Thunderbolts headquarters in the Colorado Rockies[425]. Khn'nr easily incapacitated Thunderbolts team, but before he could slay them he was talked down by their leader Norman Osborn[426][431][430]. A brilliant manipulator, Osborn convinced Khn'nr to give up on his Skrull orders and embrace the honour of the hero he was conditioned to imitate, prompting Khn'nr to fly off into space. Prior to the attack, the Thunderbolt's member Swordsman was provided with a clone of his sister Andrea[431], suspecting that she was a Skrull, Osborn ordered his troops to be prepared to kill her if she acted against them. Osborn then took the impending Skrull attack to boost the Thunderbolt's profile while discrediting Tony Stark/Iron Man and ordered his Thunderbolts to Washington D.C. to fight the Skrull invasion there.[432]

Bermuda Triangle

The Skrull virus also knocked out the primary SHIELD helicarrier[425] sending it crashing into the Bermuda Triangle. There the Skrull posing as Jarvis approached who he thought was SHIELD director Maria Hill, and ordered her to surrender[426][428]. However at the advice of Nick Fury, months earlier[315], Hill had placed a Life-Model Decoy in her place and instead of surrendering detonated the SHIELD helicarrier[430]. However the Skrull posing as Jarvis survived and managed to return to New York City.[433]

Camp Hammond

Just prior to the invasion newly appointed leader of the Hawaii based Point Men Initiative Team, the new 3-D Man uncovered that his teammate Magnitude was a Skrull and slew him. He then had himself teleported to Camp Hammond to warn his superiors. However overhearing 3-D Man trying to raise the alarm of a Skrull invasion, the Crusader—fearing being exposed and lumped in with the invaders—used his Cosmic Ring to invert the ability for 3-D Man's goggles to reveal hidden Skrulls, making it look like everybody at the 50 State Initiative was a Skrull[390]. Needing to find allies elsewhere, 3-D Man attempted to flee getting far enough for Crusader's Cosmic Ring to stop effecting his goggles. However at this point the Skrull virus hit all Skrull tech causing 3-D Man's ship to crash land in New Mexico. The virus almost felled War Machine however a fail safe using old Stane International technology saved War Machines life. By that point reports of an attack on New York by Skrull forces. "Yellowjacket" then ordered all Initiative recruits to head into battle, all complied except for the cowardly Ant-Man shrunk down to size to avoid going along[434]. Having received a recorded message from Iron Man himself sent War Machine off to a secret cache of unaffected weapons to use against the Skrull invasion, leaving mostly support staff behind. With most of his opposition gone, "Yellowjacket" then called in Skrull agents to secure Camp Hammond, unaware that they were revealing themselves and their plans to turn the camp into the heart of the "new" Skrull empire to Ant-Man.[411]

With Skrull ships landing at Camp Hammond, Bengal and other Initiative released the unstable Shadow Initiative to stop the take over of their headquarters, teaming up with Ant-Man in the process. As the Shadow Initiative were preparing for an attack, Veranke arrived at Camp Hammond from her ordeal in the Savage Land. However the Skrulls were fully aware that the Shadow Initiative were active on the base and foiled their attempt to assassinate Veranke. However Ant-Man remained free and uncaptured[415]. The Skrulls posing as "Dum Dum Dugan" and "Yellowjacket" used Camp Hammond to send out a mass message to the human race telling them to surrender to their cause, while also monitoring their efforts to compromise the various Initiative teams that had been infiltrated by Skrull operatives. Operations that were quickly crumbling thanks to the efforts of the Skrull Kill Krew who had teamed up with 3-D Man and other Initiative recruits[413]. With their plans failing, the Skrulls at Camp Hammond decided to initiate their last resort, a weapon that utilized the Negative Zone transport towers of at least three Initiative bases to create a portal that would destroy the entire United States.[412]

New York City

While on a speaking engagement in Vancouver, British Columbia, the Invisible Woman was ambushed by Skrull operatives and she was replaced by Lyja[435]. Lyja then travelled to New York City where she gained access to the Baxter Building headquarters of the Fantastic Four where she managed to fool the Thing and Human Torch and the Richards children long enough to activate a device that shunted the upper levels of the Baxter Building into the Negative Zone[425][435]. Elsewhere in the city, young Mackenzie Walden learned that her brother had been replaced by a Skrull invader.[436]

With both the Avengers and the Fantastic Four incapacitated the Skrulls launched their attack on New York City[426][436][437]. In New York at the time were the Runaways, when their teammate Xavin confronted the Skrulls and learned what was going on. When he learned that the Skrull team they have encountered had come to specifically slay Dorrek VIII, Xavin pulled his fellow Runaways out of the battle and went off to warn the Young Avengers[310]. At this point the Young Avengers were already engaged in battle against the Skrull squad that was deployed in Times Square[428]. Xavin arrived at the scene of the battle and pulled Hulkling away from the battle, but they were pursed by Chrell[310]. With the help of Young Avengers Speed and Wican and the other Runaways helped keep Hulkling safe from Chrell and his assassin X'iv[340]. The two teams then succeeded in slaying Chrell and X'iv[341]. Hulking and his teammates returned to the battle in Times Square, where the battle continued to go poorly until the timely arrival of the Initiative recruits and Nick Fury's Secret Warriors[428][411]. However the tide soon reversed and the Skrulls were gaining the upper hand on sheer numbers alone, just when Ms. Marvel arrived on the scene. Fury pulled back the Initiative recruits and Young Avengers away. Suspecting that Ms. Marvel was a Skrull, Fury blasted her with is weapon before teleporting away, leaving her at the mercy of the Skrulls[429][438][439]. At Fury's hideout, the conflicted Crusader decided once and for all that he would fight along side the heroes of Earth and defend his adopted home.[415]

Despite being left alone Ms. Marvel managed to fight off against the Skrulls. Facing a superior foe, the Skrulls then tried to disguise themselves as Earth civilians caught in the middle of the conflict. Despite her rough means, she managed to expose the Skrulls by blasting the crowd with a low power energy blast.[440] After fighting off the remaining Skrulls on in Times Square, she helped S.H.I.E.L.D. agent Hui Lin evacuate citizens to the Raft prison.[441] However she found the Raft under attack by a Skrull called Hydra Prime 001 who tortured and murdered the guards and some of the inmates. She had the immensely powerful Skrull creature follow after her and they fought all over the city. However when the Hydra Prime was brought before his fellow Skrulls, Ms. Marvel realized that they were frightened of the creature. After Hydra Prime slew all the Skrulls in the vicinity she then was able to overpower and slay the creature herself.[442]

Elsewhere in the city the novice heroine known as Jackpot clashed with a Skrull seeking to replace Spider-Man, slaying the creature after a battle across the city.[443][444][445] In addition, Front Line writer Ben Urich was caught in downtown Manhattan during the invasion. From there he recorded the personal triumphs and tragedies of every day people in the face of the Skrull invasion.[446][447][448][449] While Mackenzie Walden and her family continued to try and survive the invasion.[450][451][452][453] The invasion also freed the Punisher from prison, and seeing the chaos he rode into New York City to lay waste to every Skrull he came across.[454] The Punisher's shooting spree attracted the attention of a Super-Skrull with the powers of Hammerhead and the Kingpin, S.H.I.E.L.D. agent G.W. Bridge and his former ally Rampage. However the three humans put aside their differences long enough to take down the Skrulls attacking them, however Rampage was horribly maimed in the process.[455] The New Warriors also briefly fought the Skrulls in New York.[456]

In Space

One of the few survivors of the Skrull attack that destroyed the SWORD satellite was it's director Abigail Brand thanks to her uniform's limited life support systems[425][427]. Abigail managed to float toward a Skrull ship and force her way inside where she learned what was happening on Earth and that the ship she managed to board was also holding Reed Richards prisoner[429]. Using her knowledge of the Skrull language, Brand managed to fool the Skrulls on board into thinking she was one of their infiltrators and slew them all, then freed Reed Richards. Richards then prepared a device that would help them reveal disguised Skrulls and the pair returned to Earth. As they fled to Earth, the invasion ship was attacked and destroyed by the Skrull posing as "Captain Marvel".[430]

The Inhumans were also attacked on the moon by a Skrull invasion fleet, while somewhere deep in space a Skrull scientists attempted surgery on the real Black Bolt to channel his destructive voice into their warship in order to give it a devastating weapon[410]. When Black Bolt refused to use his powers the Skrulls then threatened to harm his son Ahura forcing him to comply. Meanwhile, the Inhuman royal family managed to stave off the Skrull invasion of their home and then left for space to search for their missing leader[22]. The royal family then approached the Kree, with Medusa pledging that the Inhumans will begin following their intended purpose of battling the Skrulls on behalf of the Kree. They then set about to obtain various items that Karnak needed to hack into the Skrulls data bases and learn the location of their leader[398]. Succeeding in their missions, the Inhumans were able to locate the Skrull ship and attacked it. During the confusion, Black Bolt managed to break free and slew his captors. The reunited Inhuman family then returned back to their home on the moon after destroying the Skrull ship.[457]

Meanwhile, War Machine followed Tony Stark's signal into space where he came across a heavily weaponed satellite piloted by his former Force Works teammate Suzie Endo.[458] The Skrull invasion fleet over Earth was then attacked by War Machine and Endo decimating many of the ships in the process.[459]

On the planet Dendron the hero Nova was attacked by a team of Super-Skrulls and was assisted by Kl'rt the original Super-Skrull. Learning of the Skrull invasion of Earth, Nova headed toward Earth with Kl'rt with him. There they were attacked by a Skrull fleet where Nova was seemingly betrayed by Kl'rt.[460] However this was a ruse on the Super-Skrull's part in order to allow both of them to escape to Earth.[461]

Not all Skrulls in space wanted to take part in the invasion of Earth. A group of Skrull fled to Knowhere and were under the protection of Knowhere's security chief Cosmo in order to escape those pursuing them for assassination.[462] However a Super-Skrull suicide bomber came to Knowhere and blew himself up in an attempt to expose the defectors, alerting the Guardians of the Galaxy that their home base was infiltrated by Skrulls.[463] After a clash with the Galactic Council and their enforcers the Luminals,[464] the Skrulls were exposed but were allowed to live in peace when the truth was learned.[462]

Washington D.C.

The Thunderbolts answered the request for assistance in Washington D.C., which was also under siege by the Skrull invaders. However the Skrulls were ready to deal with the group of "reformed" super-villains, by using Super-Skrull modelled after former members of the team, appealing to Moonstone's desire for power, Bullseye's capacity for murder, Penance's guilt over the death of hundreds of people in Stamford, Connecticut, overloading Radioactive Man to reach critical mass, and Norma Osborn's hatred of Spider-Man against the group.[432]

However despite their duplicitous ways and deceptions, the Thunderbolts managed to pull together to fight back against the Skrulls, slaying every attacker and saving the capital.[465] The last surviving Skrulls attempted one last suicide run by consuming pills that caused them to melt and merge into a massive liquid form, but this too was stopped by the Thunderbolts, winning them national praise. With Washington D.C. now safe, Osborn ordered his minions to New York to fight off the invasion happening there.[466]

San Francisco

A group of normal Skrulls were dispatched to San Francisco, unaware that it was under protection from the X-Men. The X-Men succeeded in their initial attack forced back the opening salvo. During the attack, the X-Man Nightcrawler obtained an orb that contained artificial intelligence which tried to convert the mutant hero into a loyal follower of the Skrull faith.[467] The Skrulls came back by using their own telepaths to jam the X-Men's ability to communicate via long range telepathy and then attacked the city again with Super-Skrulls. When the X-Men responded with a series of guerilla attacks the Skrulls then began kidnapping the people of San Francisco and trapping them in various towers[468]. The Skrulls then gave the X-Men an ultimatum: stand down within 24 hours and surrender or all the humans they have captured will be slaughtered. However, the X-Men were given an idea when Nightcrawler showed Beast the orb that was trying to "convert" him and explained that it was doing so because by the Skrull's logic, mutants and Skrulls were not all that different.[469] With this inspiration, Beast examined one of the dead Skrulls they captured and learned Skrull DNA was very similar to mutants. The X-Men then manufactured a strain of Stryfe's Legacy Virus which was engineered to be lethal to Skrulls only. Then under the auspices of surrender, the X-Men infected the entire Skrull armada. When the Skrulls began falling ill to the deadly virus, X-Men leader Cyclops offered them the antidote in exchange for a cure, however the Skrulls refused and flew up into the sky setting their ships to self destruct, choosing death with honour than defeat.[470]

Broxon, Oklahoma/Asgard

Beta Ray Bill managed to break free from Skrull captivity just moments prior to the invasion and return to Earth. He crashed in Asgard which at that time floated high above the town of Broxon, Oklahoma. He warned the Asgardians of the coming invasion and that the Skrulls planned to utilize his hammer Stormbreaker as a weapon. With duties needing to be carried out in the town of Broxon in his alter-ego of Dr. Donald Blake, Thor gave Beta Ray Bill his hammer Mjolnir to lead the battle against the Skrulls invading Asgard[422]. While Blake helped a woman in Broxon give birth, Beta Ray Bill led the warriors of Asgard against the Godkiller Skrulls made specifically for fighting gods. When the woman in Broxon gave birth, a Godkiller armed with weapons fashioned from Stormbreaker flew to the city intending to destroy it. Beta Ray Bill went after it and was felled trying to defend the town[471]. With the combined efforts of Thor and the Warriors Three, the heroes managed to pull Stormbreaker from the Godkiller and return it to Beta Ray Bill. Thor and Beta Ray then battled the Godkiller Skrull who was slain when Thor commanded Asgard to fall upon it, crushing the attacker to death. With the Godkiller dead, the Skrull invasion of Asgard was easily repelled.[472]

Across America

Arriving on Earth, Nova learned from his family that his brother Robert was now working at Project PEGASUS which was under attack from Skrull forces. Nova assisted Darkhawk and the newly resurrected Quasar defend Project PEGASUS from Skrulls trying to infiltrate the research lab in order to gain access to alien weapons[461]. The heroes gained the upper hand when Robert and his co-workers were able to reboot Nova's Xandarian Worldmind who intern activated the Death's Head robots that fought off the Skrulls attacking PEGASUS and their ship above.[473] Meanwhile in Minneapolis, She-Hulk and Jazinda clashed with Skrull invasion forces there. Trying to head up north to Canada to find Jazinda's ship to contact the Skrulls and inform them that they have Nogor as their prisoner, the pair were attacked by the Kl'rt the Super-Skrull. Kl'rt had come not only to liberate Nogor, but to slay his daughter Jazinda for bringing dishonour to her family.[474] However despite his efforts to slay his own daughter, She-Hulk convinced Kl'rt that he was only really mad about his son being dead and the Super-Skrull gave up his fight and left with Nogor.[391]

In the Appalachian Mountains, the New Warriors and Counter Force fought Skrulls who were attacking one of the last remaining SHIELD helicarriers, saving the crew on board. They had come to determine if the corpse of deceased members Night Thrasher (Dwayne Taylor), Namorita and Microbe in light of the Skrull invasion, their current leader the new Night Threasher believed that his brother might had been replaced by a Skrull during the Stamford Disaster that caused the super-hero Civil War months earlier. However they determined that the corpses were that of their dead teammates and gave them a proper burial.[475]

After his plane had crashed in New Mexico, 3-D Man was going to be rescued by Maverick's member "She-Thing", however he recognized the Skrull impostor, but before she could attack "She-Thing" was killed by Ryder and Riot the last surviving members of the Skrull Kill Krew. With the assistance of Charlie Chandler and his family, the 3-D Man joined the Skrull Kill Krew and joined them on a cross-country mission to destroy all the Skrull infiltrators within the Initiative. Their first stop was to slay Blacksmith of the Desert Stars[411]. With each exposition of a Skrull infiltrator the various members of the each Initiative team then joined up with the Kill Krew. They then began clearing out the Skrulls hiding out in the states of Nevada, Utah[415], Montana, Philadelphia, Georgia[413], Arkansas, Kentucky, Wisconsin, Florida, Washington, and Texas. Learning that the Skrulls were attempting to use the Negative Zone receiving towers as a doomsday weapon to destroy America, Initiative members Whiz Kid and Spinner were sent to destroy the receiving towers using their super-speed abilities.[412]

In Atlanta, the Skrulls attacked a baseball game and were opposed by the mercenary known as Deadpool. Deadpool managed to cripple the Skrull forces but to their surprise offered to join their cause instead.[476] Deadpool then allowed them to experiment on him and create Super-Skrulls that could replicate Deadpool's healing factor making essentially unkillable Skrull warriors. However Deadpool began training them and instead of being useful warriors they became mostly insane. This however was all part of a plan as Deadpool was working to steal intel from the Skrulls for Nick Fury.[477] Deadpool then fought off another Super-Skrull, and the Skrulls eventually realized how they had been tricked: when duplicating Deadpool's healing factor they forgot to take into account the reason why his healing works is due to the cancer that has infected his system. Without the cancer as a catalyst to keep the healing factor under control, the Skrulls endowed with Deadpool's healing factor began to grow into horribly deformed Skrulls unabel to move. Deadpool then killed the other Skrulls and began to transmit the data to Fury, however it was intercepted by Norman Osborn.[478]

At the Cube facility, Noh-Varr the Marvel Boy managed to take over the facility, however when the Skrull virus infected the Cube, he found Skrull infiltrators inside.[427] Breaking out of the prison, Noh-Varr crossed paths with the mortally wounded "Captain Marvel" who revealing his Skrullian origins convinced Noh-Varr to fight for the people of Earth.[479]

In the state of Oregon, Skrulls initially had the upper hand with little to no super-human interference until they were shut down by the Agents of Atlas[427]


Skrull commander K'vvvr was charged with attacking the African nation of Wakanda and destroy the Vibranium there. However his infiltrators were almost all discovered by Black Panther and his wife Storm, the rulers of Wakanda. Then using Wakandan technology, the Black Panther crippled the Skrull invasion fleet and their energy weapons forcing them to fight in hand to hand combat. Black Panther then led his people in battle against the Skrulls.[480] The Black Panther and Storm held their own against the Super-Skrull operatives while the warriors of Wakanda dealt with the regular invaders. However a lone Skrull infiltrator escaped discovery and was able to capture both the Panther and his queen.[481] However this was all a ruse as the Black Panther and Storm had really used Wakandan technology to make two captured Skrulls look like them. When K'vvvr realized this during torture he was too late to stop the Panther and Storm, who slew him and the rest of his army soon fell at the hands of the people of Wakanda. The Black Panther then had all the Skrull bodies put aboard the Skrull ship and sent it back out into space. Painted on one of the walls in the command centre in blood was the warning "This is what happens when you invade Wakanda" as a warning to other would be invaders from beyond the stars.[482]


In Russia, the Skrulls attempted to take control of the nuclear weapons found in that country in the hopes of using them against the human race. They found opposition from the Winter Guard. War Machine, after destroying Skrull ships orbiting the Earth, came to the Winter Guard's aid but his assistance was refused. Ignoring demands to leave Russian soil, War Machine fought against Skrull forces but was soon captured and found himself on a Skrull examination table.[459] With his life support failing, War Machine managed to break free and fight his way to his armor, saving his life. Destroying the Skrull ship, War Machine then returned to help Winter Guard in the battle once again. Facing defeat, one of the Skrulls attempted to overload his powers to destroy the area and set off the nuclear weapons, however War Machine grabbed the Skrull and flew him high into the air where exploded harmlessly.[483]

London and Otherworld

At some point prior to the Skrull invasion, Joint Intelligence Committee of the United Kingdom director Mortimer Grimsdale was replaced with a Skrull. When the invasion kicked off this infiltrator used his position to slay all of the Skrull Beatles except for Skrull John who was rescued by Pete Wisdom. They then joined heroes Captain Britain, Black Knight, Spitfire and the newly empowered Faiza Hussain in trying to stop the Skrull invaders from destroying the Siege Perilous the doorway into the mystical realm of Otherworld. They failed and Captain Britain was seemingly slain trying to stop the attack[484]. With the Siege Perilous destroyed, the Skrulls—whose mission it was to take over all magic—used their own portals to gain access to Otherworld convinced that they would not have opposition from Britain's heroes. However Wisdom called upon the help of his estranged wife Tink the fairy to gain access to Otherworld. There they helped Tink's father Oberon defend Otherworld from the Skrulls. However they were unable to stop the Skrulls from slaying the Lady of the Lake and the Green Knight and gather the mystical artifacts they needed to create a Super-Skrull that was entirely magic based to destroy London.[485]

While Britain's heroes defended London, Pete Wisdom was drawn to a portal where the wizard Merlin the Magician was trapped after being infected by the Fury. Wisdom freed Merlin but also many demons that were trapped there as well. Merlin then resurrected Captain Britain and armed him with the sword Excalibur to defend London[486]. While Captain Britain fought against the mystical Super-Skrull, the rest of his comrades were captured by the Skrull army and Skrull John was executed for betraying his people. Before the rest of the group could be slain they were rescued by Satannish and a horde of demons. Satannish, grateful to Wisdom for freeing him and the other demons and granted Wisdom a boon. Wisdom uttered the words "No more Skrulls" and the demon used his magics to incinerate all the Skrulls in the United Kingdom, ending the conflict there.[487]

Negative Zone

The Human Torch, Thing as well as Franklin and Valeria Richards found themselves trapped in the Negative Zone by Lyja, the Torch's estranged wife who sought revenge against Johnny for abandoning her for years[435]. While the Thing protected the children from creatures from the Negative Zone, Johnny and Lyja fought it out but soon gave up on trying to kill each other and instead rekindled their romance before they were attacked by a massive jellyfish like creature. Johnny saved Lyja's life and returned her to the ruins of the Baxter Building. There Franklin deduced that their only way out of the Negative Zone was to seek out one of the many genius villains locked up in Prison 42[251]. Going to the prison, Johnny, Lyja and the Thing freed the Tinkerer who agreed to help them return home out of sympathy for Franklin and Valeria. However before they left Lyja decided that she would stay in the Negative Zone to find a new life. After bidding the Torch farewell, the displaced heroes returned themselves to the positive matter universe.[488]

Realm of the Gods

On yet another front, the pantheon gods of Earth were at risk of being destroyed an enslaved by the Skrull deities much like other pangalatic gods across the universe. Athena then called upon the Council of Godheads to meet and address the problem. They then selected champions to travel to the realm of the Skrull gods and combat them there. The group selected the Greek god Hercules and his human ally Amadeus Cho, Snowbird of the Inua, the Eternal Ajak was chosen to represent the Apu gods, Amatsu-Mikaboshi of the Amatsu-Kami, and Atum aka the Demogorge to do battle against the Skrull gods Sl'gurt't and Kly'bn[4]. The group fought their way through Nightmare's realm unaware that Amadeus' cycote Kirby had been replaced with a Skrull[8]. When they arrived in the realm of the the Skrull gods Atum discovered that "Kirby" was a Skrull and slew it and they were soon attacked by the gods whom the Skrulls had enslaved during centuries past. The enslaved gods were vanquished thanks largely to Snowbird, who seemingly sacrificed her life transforming into Neooqtoq to destroy their attackers[489]. At first the so-called God Squad proved to be no match for the two Skrull deities. However it was not until Amadeus pretended to abandon Hercules did the Grecian put in his all into the battle. Ultimately, Sl'gurt't was slain by Amatsu-Mikaboshi and Kly'bn slain by Snowbird. The Skrulls realm was then shattered by a blow from Hercules that was so powerful every Skrull idol of worship across the universe exploded. This gathering of gods managed to strike the greatest blow against the Skrull invasion: a blow against their faith.[7]

Final Battle

In the Savage Land, Reed Richards and Abigail Brand landed in the area and Reed used his device to reveal who among the gathered heroes were Skrulls[430]. The combined Avengers teams[479], the Thunderbolts[466], members of the Initiative, and the Secret Warriors[412] joined forces for a final battle against the Skrulls in Central Park. During the final battle, Luke Cage's wife Jessica Jones left their child Denielle in the care of the Skrull Jarvis while she joined the battle in Central Park. It was also here that Criti Noll revealed herself to be a Skrull posing as Yellowjacket. As the heroes began gaining the upper hand, Noll activated a device that effected the Wasp's powers making her into a massive doomsday weapon to kill the heroes[433], however her fellow Avenger Thor was forced to slay her to save the others. In the final push the battle was suddenly one when Norman Osborn was able to fire the shot that killed Veranke. With their leader killed the Skrulls were easily defeated and the remainder of their fleet in space destroyed. The heroes of Earth also found a ship containing all their detained comrades. When Jarvis was found among the captured, Jessica Jones and Luke Cage rushed back to Avengers Tower to find that the Skrull posing as Jarvis had disappeared with their child[433]. Meanwhile, Criti Noll survived the battle and attempted to use super-speed to return to Camp Hammond. He was followed initially by the Crusader and then by the rest of the Initiative and the Skrull Kill Krew. While the Crusader slew Criti Noll, 3-D Man eliminated the Skrull posing as Dum Dum Dugan. In the aftermath of the battle, 3-D Man suddenly saw the Crusader for the Skrull that he was—and not realizing that Z'Reg was loyal to Earth—shot him dead. Dying Z'Reg wished that things happened differently and was teleported away by his ring [412]. While a lot of Skrulls were captured as prisoners of war[490], many ended up being captured or managed to escape.

Interestingly enough, the Skrull/Cow hybrids that had been breeding for years had reached enough of a population and awareness that they were known to the United States government. These Skrulls had managed to integrate into nearly every aspect of Earth society, particular the media. For immunity and to keep their existence a secret, the Earth born Skrulls agreed to boost support against the alien Skrull invaders through the media.[491]


Following the defeat of the Skrulls, the head of homeland security was appointed to Norman Osborn, who was now a hero for killing Veranke. He then disbanded SHIELD and created his own organization called HAMMER, many of the surviving Skrulls who were captured were handed over to this organization for questioning. One Skrull prisoner was literally fed to Venom by Osborn[492]. Immediately after the the invasion, Luke Cage and his fellow New Avengers began looking for the Skrull who posed as Jarvis in order to find his daughter. Although they managed to find a Skrull hiding among a number of former S.H.I.E.L.D. agents, they were unable to learn the location of "Jarvis", forcing Cage to make a deal with Norman Osborn[493]. Cage was given access to one of the Skrull prisoners who was tortured into revealing the Skrull invaders secret warehouse meeting place, the most likely spot where "Jarvis" could have went to get off the planet, however he could only do so with someone operating the teleportation system for him. Cage went to the warehouse and was able to find "Jarvis" and his daughter. Luke then agreed to help the Skrull return home in exchange for his daughter. However when "Jarvis" handed the baby over he was shot in the head by Bullseye who was hiding in a nearby building with a sniper rifle [494]. Norman Osborn had imprisoned Elektra to learn why the Skrulls experimented and tortured her when their other captives were relatively unharmed. However before he could succeed in learning this information Elektra escaped captivity[495]. Osborn then pulled two captive Skrulls for information, torturing them in order to learn that the Skrulls were trying to learn Elektra's ability to resurrect herself after death.[308] He then continued to torture them to learn when Elektra was replaced, and eventually murdered the prisoners in front of H.A.M.M.E.R. accountants to show them the price of disobeying his wishes.[496]

Meanwhile in space, a fleet of Skrulls trying to escape Earth were pursued by the Inhumans into space. They were followed into the Arkano Nebula where they the fleet was wiped out by Black Bolt using a weapon that amplified his destructive voice[497][498]. Following the defeat of their empire a group of Skrulls decided to become pacifists and found a new would-be god in Beta Ray Bill. Bill attempted to help them find shelter from S'Kaan a surviving Skrull Godkiller. Beta Ray Bill defeated S'Kaan but his life was spared. The Skrullian Monks then healed him and convinced that Beta Ray Bill was not their new god, but a agent of this force they healed S'Kaan who assisted them in setting up a new home aboard the Remnants starbase[499]. Many slain Skrulls became the power source for the Kree Babel Spire defence system.[500]

Back on Earth, Ryder of the Skrull Kill Krew discovered the Skrull Cows hiding in society and became devoted to hunting them down and destroying them all[501]. With his teammates slowly coming back to life, Ryder sought out the aid of HAMMER scientist Viola Reichardt to examine his DNA. She learned that the Kill Krew members were not only coming back to life, but becoming Skrulls themselves. Meanwhile, Ryder went to Los Angeles where he found an entire group of young Cow-Skrulls who used a night club to lure humans to slaughter, killing them all.[115] Ryder and the resurrected Riot then went after another Skrull nightclub called the Reverse Rodeo where Skrulls rode human beings like cattle in a faux-rodeo.[502] As the rest of the Krew was coming back to life, Riot soon soon began dating a Skrull she met. The Krew was then manipulated by Moonstomp into attacking a restaurant full of Skrulls where Riot and her date were. They were assisted by a Skrull who was posing as Wolverine.[503] After the Krew slaughtered Skrulls on the Stampede web series, the Cow-Skrulls sent an entire army to their home for revenge, but they too were slaughtered by the Krew.[504] The crew slaughtered another restaurant full of Skrulls, prompting the Cow-Skrull elite to press their influence to get H.A.M.M.E.R. and Norman Osborn to act. Osborn then called a meeting with the Skrulls the Kill Krew appeared again and slew the entire group of cow Skrulls before going into hiding again.[491]

Another Skrull who evaded capture during the invasion was Koru Kaviti who fled to the nation of Madripoor with another Skrull who posed as Spider-Man. SWORD agent Abigail Brand recruited Spider-Woman to eliminate Kavitti. Upon arriving in Madripoor, Spider-Woman was attacked by the Skrull posing as Spider-Man and managed to slay it.[505] Later on her mission, Spider-Woman was approached and offered a position in Hydra again by Madam Hydra. As a gift, Madam Hydra allowed Spider-Woman to do what she saw fit with a Skrull that Hydra managed to capture during the Skrull invasion. When the Skrull refused to talk it tried to attack her and she was forced to slay it.[506] Ultimately, Spider-Woman managed to track Koru to a local bar and after an epic battle managed to slay the Skrull with the help of her fellow New Avengers.[507] Back in the United States the mutant terrorist Mystique discovered an abandoned Super-Skrull facility where a number of failed attempts to create a Skrull impostor of Captain Marvel. She released one of these Mar-Vell Super-Skrulls an unleashed him on Ms. Marvel in Seattle. This Skrull violently exploded in the skies above Washington State when his powers overloaded.[508] A Skrull who survived the battle in the Savage Land accidentally set off a S.H.I.E.L.D. alert while working in the area bringing Mockingbird and the new Dominic Fortune to the region. Exposing the phony Skrull, Mockingbird burned their bodies.[509]

Klobok soon resurfaced and used his arcane knowledge to try and re-establish a bond with the estranged Dire Wraiths in an attempt to reform the Skrull empire[116]. However by this time the Dire Wraiths had been banished to Limbo, and so Klobok made a mystical spell to bring the Wraithworld from Limbo. This threatened to destroy the planet Galador home of the Spaceknights. A group of intergalactic heroes calling themselves the Annihilators came to Galadors aid, unaware that Klobok had infiltrated them by posing as Doctor Dredd[510]. Although Wraithworld was brought back to Earth, Klobok was slain in battle and the Wraiths made peace with their longtime foes the Galadorians[116]. Back on Earth, former Avengers Wonder Man gathered a group of super-humans to fight his former comrades called the Revengers. Among the groups members was Virtue who returned to costumed heroics after being a faith healer.[511][512]

Another group of Skrulls who fled the losing invasion resurfaced and tried to return home. Manipulating the young X-Man known as Pixie they managed to break into the Baxter Building and Horizon Labs to gain access to a hidden Skrull ship. However their leader Nazum sought to get revenge against Earth and unleashed Skrull war machines.[513] While the X-Men and the Future Foundation fought the devices, Pixie and Spider-Man worked together to stop the Skrulls whom Pixie allowed to leave without further harm. The Skrull deserted left Nazum on a uninhabited world.[514] A number of Skrull corpses that were taken during the invasion ended up in the custody of a newly reformed SHIELD. These corpses were stolen by Madame Hydra when she raided a SHIELD warehouse. Their possible use is unknown at this time.[515]

Not long after a group of Skrulls returned to Earth and captured Captain America and Cyclops, who at the time were bitter enemies. Commander K'thron warned them that Cadre K had returned to Earth and were planning another Skrull invasion[516]. The pair put aside their differences the pair used SHIELD resources to learn that a surviving Skrull invader was captured by Dr. Doom and was being held captive in Latveria. The pair fought their way into Castle Doomstadt. Doom eventually allowed them to take the captured Skrull, but refused to let it leave alive[517]. However this Skrull was faking it and upon bringing him back to the United States, the Skrull led the two heroes to a Brysons Bison Burgers chain that—with the help of Ant-Man, Emma Frost and the Stepford Cuckoos—discovered that all the staff were Skrulls. Stranger still, the bison beef served at the restaurant was all based on mutant Skrull DNA[518]. They tracked the source of the beef to Sagittariuan Farms in South Dakota. There they discovered that Cadre K had taken over the ranch and used it as a front to collect the last of the Cow-Skrull hybrids in order to prevent them from being slaughtered and prevent the human food supply from being tainted by DNA altering Skrull meet[519]. Realizing Cadre K's altruistic intents, the Avengers and Cyclops's X-Men defended them from Dr. Doom[520], SHIELD agents and the Skrulls that sent Cyclops and Captain America on their mission to begin with. The duo managed to convince all three parties that Cadre K posed no threat to the world.[521]


Following the invasion, the Skrull empire was broken up and ruled by different warlords which included Kl'rt, My'rl, Dm'yr, Gy'pl.[522] When the Builders, ancient aliens who claim to have created all existence, deemed that Earth should be destroyed, they sent a massive invasion fleet to attack the Earth, destroying various worlds as they passed.[523] This swath of destruction also included the Skrull world Hy'lt Minor, slaying two-hundred billion Skrulls including warlord Dm'yr.[522] A group of Skrulls fled to Earth and managed to slip through S.W.O.R.D. defences.[523] However they were quickly apprehended by the Avengers.[524]

Kl'rt was selected as the warrior to represent the entire Skrull empire when they went to the planet Nomad to join the Galactic Council to join a coalition of alien races which included the Kree, the Shi'ar, the Brood, Spartax, and various other races including the Avengers. However when they set up an ambush for the Builders in a sector of space called the Corridor the Council was overpowered and they were forced to retreat.[522] They regrouped over the Skrull agricultural world of Whaan Prime, but the Builders followed them there and unleashed a plague upon the world. While the Council members tried to save as many of the Skrulls on the planet surface, billions still died.[525]

The Council retreated to the Behemoth Ringworld to regroup, but were forced to retreat when the ruler of the Spartax betrayed their location to the Builders the planet was attacked forcing another evacuation.[526] One hundred million refugees died in the attack, many of whom were Skrull survivors who fled previous attacks[527]. After regrouping Kl'rt sided with the other Council members to allow Captain America to take the lead in. Cap in turn suggested that they surrender.[528] However this surrender was a false front and allowed the Avengers Thor to slay the Builder who had just taken over the Kree throneworld of Hala[529]. The Skrull armies were involved in the following skirmishes that saw the Council begin to regain ground and liberate planets under the Builders' rule, but the Skrulls saw heavy casualties.[530] However many worlds were liberated and eventually the coalition, under Captain America's command, eventually beaten back the Builders and their path of destruction. Learning that Earth was now under attack by the forces of Thanos, Kl'rt lent his own warriors to help liberate the Earth, a plant that his people tried to enslave months earlier[531]. After an attack on Thanos' forces on the Saturnian moon of Titan,[532] Skrull forces attacked the armadas of Thanos in Earth's orbit. Learning that one of Thanos' agents, Black Dwarf now controlled the SWORD space station, Super-Skrull personally joined, Shi'ar leader Gladiator, Ronan the Accuser, and Annihilius in slaying him.[533] With the path to Earth cleared, the Avengers ultimately defeated Thanos and his followers. With the threat of both Thanos and the Builders over, the Skrull empire reformed with Tarnax II becoming the new throneworld and Kl'rt being crowned the new emperor of the Skrulls.[534]


  1. Thor #215
  2. Avengers Vol 5 #2
  3. 3.0 3.1 3.2 Silver Surfer Vol 3 #5
  4. 4.0 4.1 Incredible Hercules #117
  5. 5.0 5.1 Rom #50
  6. Blackwulf #9
  7. 7.0 7.1 Incredible Hercules #120
  8. 8.0 8.1 8.2 Incredible Hercules #118
  9. 9.0 9.1 She-Hulk Vol 2 #31
  10. 10.0 10.1 Avengers #133
  11. Avengers #134
  12. Captain Marvel #38
  13. X-Men Annual #11
  14. Thunderbolts #45
  15. Thunderbolts #46
  16. Captain America Annual #7
  17. 17.0 17.1 17.2 Avengers #94
  18. Europa #0
  19. Fantastic Four Vol 4 #10
  20. Marvel Westerns: Kid Colt and the Arizona Girl #1
  21. 21.0 21.1 Fantastic Four #91
  22. 22.0 22.1 Secret Invasion: Inhumans #2
  23. Blackwulf #2
  24. 24.0 24.1 Marvel: The Lost Generation #4
  25. Marvel: The Lost Generation #1
  26. Marvel Premiere #35
  27. Marvel Premiere #36
  28. 28.0 28.1 Marvel Premiere #37
  29. Marvel: The Lost Generation #2
  30. Marvel: The Lost Generation #6
  31. Marvel: The Lost Generation #7
  32. Wisdom #2
  33. 33.0 33.1 Fantastic Four #206
  34. 34.0 34.1 34.2 34.3 Avengers Annual #14
  35. Iron Man #65
  36. Amazing Adult Fantasy #7
  37. Marvel: The Lost Generation #11
  38. Originally, the story in Amazing Adult Fantasy #7 was a stand alone sci-fi tale, however the premise was used again in Marvel: The Lost Generation #11 replacing the generic alien in the story with a Skrull, updating the concepts -- from a book of alien languages to a communicator device -- and changing the bald scientist to that of Reed Richards. The Skrulls! #1 handbook confirms that the two stories are the same, as such the details of Lost Generation #11 retcon some of the differences of Amazing Adult Fantasy #7
  39. 39.0 39.1 X-Men: The Hidden Years #16
  40. Marvel: The Lost Generation #12
  41. Fantastic Four #2
  42. 42.0 42.1 42.2 Marvel Team-Up Vol 3 #20
  43. Fantastic Four #18
  44. 44.0 44.1 Fantastic Four #32
  45. 45.0 45.1 Fantastic Four #37
  46. 46.0 46.1 Thor #142
  47. X-Men First Class #6
  48. Fantastic Four Annual #3
  49. Fantastic Four #48
  50. Fantastic Four #50
  51. X-Men: First Class Giant-Size Special #1
  52. Fantastic Four #89
  53. Fantastic Four #90
  54. Fantastic Four #92
  55. Fantastic Four #93
  56. Fantastic Four #64
  57. Marvel Super-Heroes #12
  58. Captain Marvel #2
  59. Captain Marvel #3
  60. 60.0 60.1 Ms. Marvel Vol 2 #25
  61. Avengers #69
  62. Avengers #70
  63. Avengers #71
  64. Avengers #91
  65. 65.0 65.1 Avengers #93
  66. 66.0 66.1 Avengers #92
  67. 67.0 67.1 Avengers #97
  68. 68.0 68.1 Fantastic Four Annual #17
  69. 69.0 69.1 Avengers #96
  70. 70.0 70.1 Marvel Team-Up #5
  71. 71.0 71.1 Marvel Premiere #24
  72. 72.0 72.1 Young Avengers #11
  73. 73.0 73.1 Young Avengers #10
  74. New Avengers: Illuminati Vol 2 #1
  75. 75.0 75.1 75.2 New Avengers #40
  76. New Avengers #44
  77. Captain Marvel #25
  78. Captain Marvel #26
  79. Captain Marvel #27
  80. Avengers #125
  81. 81.0 81.1 Marvel Chillers #3
  82. Marvel Chillers #5
  83. Marvel Chillers #6
  84. Marvel Chillers #7
  85. 85.0 85.1 Marvel Team-Up #61
  86. Fantastic Four #173
  87. Fantastic Four #174
  88. Fantastic Four #175
  89. Marvel Team-Up #60
  90. Marvel Team-Up #62
  91. Captain Marvel #53
  92. Marvel Two-In-One #45
  93. Fantastic Four #204
  94. 94.0 94.1 94.2 Fantastic Four #205
  95. 95.0 95.1 Fantastic Four #212
  96. Nova #25
  97. Fantastic Four #208
  98. Fantastic Four #209
  99. Fantastic Four #213
  100. Fantastic Four #214
  101. Avengers #209
  102. 102.0 102.1 102.2 Mighty Avengers #14
  103. Wolverine: First Class #19
  104. Wolverine: First Class #20
  105. X-Men #136
  106. X-Men #137
  107. 107.0 107.1 107.2 Fantastic Four Annual #18
  108. Fantastic Four Annual #15
  109. Rom #24
  110. Death of Captain Marvel #1
  111. 111.0 111.1 111.2 111.3 Silver Surfer Vol 3 #14
  112. Fantastic Four #249
  113. Fantastic Four #250
  114. 114.0 114.1 114.2 Skrull Kill Krew #2
  115. 115.0 115.1 Skrull Kill Krew Vol 2 #1
  116. 116.0 116.1 116.2 Annihilators #4
  117. Fantastic Four #256
  118. Uncanny X-Men #369
  119. X-Men Vol 2 #89
  120. Uncanny X-Men #370
  121. 121.0 121.1 X-Men Vol 2 #90
  122. Fantastic Four #257
  123. Fantastic Four #244
  124. 124.0 124.1 Fantastic Four #261
  125. 125.0 125.1 X-Men Vol 2 #95
  126. 126.0 126.1 Marvel Knights: Spider-Man #16
  127. 127.0 127.1 Marvel Knights: Spider-Man #15
  128. 128.0 128.1 128.2 Silver Surfer Annual #1
  129. 129.0 129.1 129.2 Avengers #259
  130. 130.0 130.1 130.2 Silver Surfer Vol 3 #2
  131. 131.0 131.1 131.2 Silver Surfer Vol 3 #6
  132. 132.0 132.1 132.2 Fantastic Four Annual #19
  133. 133.0 133.1 133.2 Silver Surfer Vol 3 #13
  134. Rom #49
  135. 135.0 135.1 Alpha Flight #10
  136. Alpha Flight #9
  137. 137.0 137.1 137.2 Fantastic Four #358
  138. Marvel Super Heroes Secret Wars #1
  139. Marvel Super Heroes Secret Wars #12
  140. Fantastic Four #264
  141. Ljya posed as Alicia Masters from Fantastic Four #264 through 356. For more details on this deception see the profile on Lyja
  142. Namor the Sub-Mariner #25
  143. Power Man and Iron Fist #111
  144. Power Man and Iron Fist #112
  145. Power Man and Iron Fist #113
  146. 146.0 146.1 Power Man and Iron Fist #121
  147. Avengers #260
  148. 148.0 148.1 148.2 Silver Surfer Vol 3 #7
  149. Avengers #261
  150. 150.0 150.1 150.2 Silver Surfer Vol 3 #1
  151. Power Man and Iron Fist #125
  152. Namor the Sub-Mariner #22
  153. West Coast Avengers Annual #2
  154. Avengers Annual #16
  155. Silver Surfer Vol 3 #3
  156. Silver Surfer Vol 3 #4
  157. Silver Surfer Vol 3 #8
  158. Silver Surfer Vol 3 #9
  159. Silver Surfer Vol 3 #10
  160. New Mutants #92
  161. Silver Surfer Vol 3 #21
  162. Silver Surfer Vol 3 #11
  163. Silver Surfer Vol 3 #12
  164. Silver Surfer Vol 3 #20
  165. 165.0 165.1 165.2 Silver Surfer Vol 3 #28
  166. Silver Surfer Vol 3 #25
  167. Silver Surfer Vol 3 #26
  168. Silver Surfer Vol 3 #27
  169. 169.0 169.1 Incredible Hulk #375
  170. 170.0 170.1 Fantastic Four Annual #24
  171. 171.0 171.1 Incredible Hulk #419
  172. Silver Surfer Vol 3 #29
  173. Silver Surfer Vol 3 #30
  174. Silver Surfer Vol 3 #31
  175. Thor #413
  176. Avengers Spotlight #25
  177. Impossible Man Summer Vacation #1
  178. Incredible Hulk #374
  179. 179.0 179.1 Fantastic Four #349
  180. Fantastic Four #347
  181. Fantastic Four #348
  182. Spider-Man / Fantastic Four #3
  183. 183.0 183.1 Namor the Sub-Mariner #18
  184. Namor the Sub-Mariner #8
  185. Namor the Sub-Mariner #10
  186. Namor the Sub-Mariner #11
  187. 187.0 187.1 Uncanny X-Men #276
  188. 188.0 188.1 Uncanny X-Men #277
  189. Uncanny X-Men #265
  190. Uncanny X-Men #273
  191. Uncanny X-Men #274
  192. Uncanny X-Men #275
  193. Infinity Gauntlet #1
  194. Infinity Gauntlet #6
  195. Namor the Sub-Mariner #14
  196. Namor the Sub-Mariner #15
  197. Namor the Sub-Mariner #16
  198. Namor the Sub-Mariner #17
  199. Impossible Man Summer Vacation #2
  200. Fantastic Four #357
  201. 201.0 201.1 Fantastic Four #366
  202. Avengers: West Coast #80
  203. 203.0 203.1 Avengers: West Coast #82
  204. Thor #445
  205. Iron Man #279
  206. Thor #446
  207. Quasar #34
  208. Wonder Man Vol 2 #9
  209. Fantastic Four #369
  210. Fantastic Four #370
  211. 211.0 211.1 Sensational She-Hulk #46
  212. Sensational She-Hulk #44
  213. New Avengers: The Reunion #1
  214. Avengers: West Coast #90
  215. New Avengers: The Reunion #2
  216. Avengers: West Coast #91
  217. For more information on this infiltration see the entry for H'rpra
  218. New Avengers: The Reunion #3
  219. Fantastic Four #371
  220. Fantastic Four #372
  221. Fantastic Four #374
  222. For more information on Lyja's time with the Fantastic Four see Lyja, Fantastic Four
  223. Fantastic Four #377
  224. Fantastic Four #378
  225. Quasar #48
  226. Infinity Crusade #3
  227. Thor #465
  228. Infinity Crusade #6
  229. Silver Surfer Vol 3 #89
  230. Silver Surfer Vol 3 #90
  231. Fantastic Four #379
  232. Fantastic Four #383
  233. Avengers: West Coast #100
  234. Nova Vol 2 #15
  235. Nova Vol 2 #16
  236. Incredible Hulk #418
  237. Silver Surfer Vol 3 #105
  238. Force Works #1
  239. Force Works #13
  240. Force Works #14
  241. See Force Works #15Force Works #16Century: Distant Sons #1
  242. See Star Masters #1 through 3 and Cosmic Powers Unlimited #4 and 5
  243. 243.0 243.1 Skrull Kill Krew #1
  244. 244.0 244.1 Skrull Kill Krew #3
  245. Skrull Kill Krew #4
  246. Skrull Kill Krew #5
  247. Spider-Man Team-Up #2
  248. Silver Surfer: Dangerous Artifacts #1
  249. Spider-Man Human Torch #5
  250. Onslaught Marvel Universe #1
  251. 251.0 251.1 251.2 Secret Invasion: Fantastic Four #2
  252. See Fantastic Four (Vol. 2) #1 through 13, Captain America (Vol. 2) #1 through 13, Iron Man (Vol. 2) #1 through 13 and Avengers (Vol. 2) #1 through 13
  253. Heroes Reborn: The Return #4
  254. Captain America Vol 3 #3
  255. Captain America Vol 3 #4
  256. Captain America Vol 3 #5
  257. Captain America Vol 3 #6
  258. Captain America Vol 3 #7
  259. Deadpool Vol 3 #24
  260. Deadpool Vol 3 #25
  261. Nova Vol 3 #3
  262. Captain Marvel Vol 4 #8
  263. Captain Marvel Vol 4 #9
  264. Captain Marvel Vol 4 #10
  265. 265.0 265.1 Uncanny X-Men #376
  266. Wolverine Vol 2 #145
  267. See James Howlett (Skrull) (Earth-616) for more details
  268. X-Men Vol 2 #92
  269. X-Men Vol 2 #93
  270. X-Men Vol 2 #94
  271. Astonishing X-Men Vol 2 #3
  272. Uncanny X-Men #375
  273. Uncanny X-Men #377
  274. X-Men Vol 2 #97
  275. Uncanny X-Men #378
  276. Cable #11
  277. Wolverine Vol 2 #148
  278. X-Men Unlimited #26
  279. X-Men Vol 2 #98
  280. Uncanny X-Men #379
  281. Avengers: Forever #12
  282. Avengers Vol 3 #35
  283. 283.0 283.1 Maximum Security #3
  284. Maximum Security: Dangerous Planet #1
  285. Maximum Security #1
  286. Bishop the Last X-Man #15
  287. X-Men Vol 2 #107
  288. X-Men Unlimited #29
  289. Maximum Security #2
  290. Fantastic Four Vol 3 #37
  291. Infinity Abyss #2
  292. 292.0 292.1 Thing: Freakshow #3
  293. Thing: Freakshow #1
  294. Thing: Freakshow #2
  295. Thing: Freakshow #4
  296. B-Sides #1
  297. B-Sides #2
  298. B-Sides #3
  299. JLA/Avengers #1
  300. JLA/Avengers #4
  301. 301.0 301.1 Avengers #503
  302. Thor Vol 2 #85
  303. 303.0 303.1 Marvel Team-Up Vol 3 #23
  304. 304.0 304.1 Avengers: The Initiative #15
  305. Young Avengers #1
  306. For more about Hulklings activities see Dorrek VIII
  307. Secret War #5
  308. 308.0 308.1 Dark Reign: Elektra #3
  309. 309.0 309.1 New Avengers #29
  310. 310.0 310.1 310.2 Secret Invasion: Runaways/Young Avengers #1
  311. Giant-Size Spider-Woman #1
  312. New Avengers #14
  313. 313.0 313.1 313.2 313.3 313.4 New Avengers #42
  314. 314.0 314.1 New Avengers #43
  315. 315.0 315.1 Mighty Avengers #12
  316. 316.0 316.1 Mighty Avengers #16
  317. New Avengers #1
  318. New Avengers #2
  319. New Avengers #3
  320. New Avengers #4
  321. 321.0 321.1 New Avengers #41
  322. New Avengers #6
  323. See Veranke for further details of her various battles with the New Avengers
  324. Marvel Team-Up Vol 3 #10
  325. Marvel Team-Up Vol 3 #2
  326. Marvel Team-Up Vol 3 #4
  327. Marvel Team-Up Vol 3 #5
  328. Marvel Team-Up Vol 3 #8
  329. Marvel Team-Up Vol 3 #9
  330. Marvel Team-Up Vol 3 #11
  331. Marvel Team-Up Vol 3 #13
  332. Young Avengers #9
  333. Young Avengers #12
  334. Marvel Team-Up Vol 3 #24
  335. Marvel Team-Up Vol 3 #25
  336. Marvel Knights: Spider-Man #14
  337. Marvel Knights: Spider-Man #17
  338. Marvel Knights: Spider-Man #18
  339. 339.0 339.1 Runaways Vol 2 #7
  340. 340.0 340.1 Secret Invasion: Runaways/Young Avengers #2
  341. 341.0 341.1 Secret Invasion: Runaways/Young Avengers #3
  342. Runaways Vol 2 #8
  343. Runaways Vol 2 #17
  344. For more on Xavin's adventures with the Runaways refer to Xavin (Earth-616)
  345. Drax the Destroyer #1
  346. Drax the Destroyer #2
  347. Drax the Destroyer #3
  348. Drax the Destroyer #4
  349. House of M #1
  350. 350.0 350.1 New Avengers #45
  351. House of M #8
  352. Uncanny X-Men #466
  353. For more on Flaw's activities see Flaw (Earth-616) and Shi'ar Death Commandos (Earth-616)
  354. Uncanny X-Men #478
  355. Doc Samson Vol 2 #1
  356. Annihilation: Prologue #1
  357. 357.0 357.1 Annihilation #2
  358. Annihilation: Silver Surfer #1
  359. Annihilation: Silver Surfer #2
  360. Annihilation: Silver Surfer #3
  361. Annihilation: Silver Surfer #4
  362. Annihilation: Super-Skrull #1
  363. Annihilation: Super-Skrull #2
  364. Annihilation: Super-Skrull #3
  365. Annihilation: Super-Skrull #4
  366. 366.0 366.1 Annihilation #1
  367. Annihilation: Ronan #2
  368. Annihilation: Ronan #3
  369. Annihilation: Ronan #4
  370. Annihilation #3
  371. Annihilation #4
  372. Annihilation #5
  373. Annihilation #6
  374. Annihilation: Conquest Prologue #1
  375. See Super-Skrull for more details on his involvement in the Kree Empire
  376. Annihilation: Heralds of Galactus #1
  377. Incredible Hulk Vol 2 #91
  378. Civil War #1
  379. 379.0 379.1 Captain Marvel Vol 6 #5
  380. Civil War: The Return #1
  381. Civil War #7
  382. Captain America Vol 5 #25
  383. Secret Invasion Prologue #1
  384. Mighty Avengers #17
  385. Wisdom #1
  386. See John the Skrull for more detail on specific missions
  387. Wisdom #6
  388. 388.0 388.1 Avengers: The Initiative #1
  389. Mighty Avengers #1
  390. 390.0 390.1 390.2 390.3 Avengers: The Initiative #14
  391. 391.0 391.1 She-Hulk Vol 2 #33
  392. She-Hulk Vol 2 #22
  393. See Jazinda for more details
  394. 394.0 394.1 Black Panther Vol 4 #33
  395. Black Panther Vol 4 #31
  396. Black Panther Vol 4 #32
  397. Silent War #6
  398. 398.0 398.1 398.2 Secret Invasion: Inhumans #3
  399. Incredible Hulk Vol 2 #105
  400. World War Hulk #1
  401. World War Hulk #5
  402. New Avengers #46
  403. New Avengers #27
  404. New Avengers #28
  405. New Avengers #30
  406. New Avengers #31
  407. New Avengers #32
  408. Mighty Avengers #7
  409. New Avengers: Illuminati Vol 2 #5
  410. 410.0 410.1 Secret Invasion: Inhumans #1
  411. 411.0 411.1 411.2 411.3 411.4 Avengers: The Initiative #16
  412. 412.0 412.1 412.2 412.3 412.4 412.5 412.6 412.7 412.8 Avengers: The Initiative #19
  413. 413.0 413.1 413.2 413.3 Avengers: The Initiative #18
  414. Avengers: The Initiative #7
  415. 415.0 415.1 415.2 415.3 Avengers: The Initiative #17
  416. Avengers: The Initiative Annual #1
  417. Avengers: The Initiative #8
  418. Captain Marvel Vol 6 #3
  419. Captain Marvel Vol 6 #4
  420. Ms. Marvel Vol 2 #26
  421. Ms. Marvel Vol 2 #27
  422. 422.0 422.1 Secret Invasion: Thor #1
  423. X-Factor Vol 3 #33
  424. X-Factor Vol 3 #34
  425. 425.0 425.1 425.2 425.3 425.4 425.5 Secret Invasion #1
  426. 426.0 426.1 426.2 426.3 Secret Invasion #2
  427. 427.0 427.1 427.2 427.3 427.4 Secret Invasion: Who Do You Trust? #1
  428. 428.0 428.1 428.2 428.3 Secret Invasion #3
  429. 429.0 429.1 429.2 Secret Invasion #4
  430. 430.0 430.1 430.2 430.3 430.4 Secret Invasion #5
  431. 431.0 431.1 Thunderbolts #122
  432. 432.0 432.1 Thunderbolts #123
  433. 433.0 433.1 433.2 Secret Invasion #7
  434. Avengers: The Initiative #5
  435. 435.0 435.1 435.2 Secret Invasion: Fantastic Four #1
  436. 436.0 436.1 Secret Invasion: Home Invasion #1
  437. Secret Invasion: Home Invasion #2
  438. Secret Invasion: Home Invasion #3
  439. Secret Invasion: Home Invasion #4
  440. Ms. Marvel Vol 2 #28
  441. Ms. Marvel Vol 2 #29
  442. Ms. Marvel Vol 2 #30
  443. Secret Invasion: The Amazing Spider-Man #1
  444. Secret Invasion: The Amazing Spider-Man #2
  445. Secret Invasion: The Amazing Spider-Man #3
  446. Secret Invasion: Front Line #1
  447. Secret Invasion: Front Line #2
  448. Secret Invasion: Front Line #3
  449. Secret Invasion: Front Line #4
  450. Secret Invasion: Home Invasion #5
  451. Secret Invasion: Home Invasion #6
  452. Secret Invasion: Home Invasion #7
  453. Secret Invasion: Home Invasion #8
  454. Punisher War Journal Vol 2 #24
  455. Punisher War Journal Vol 2 #25
  456. New Warriors Vol 4 #14
  457. Secret Invasion: Inhumans #4
  458. Iron Man: Director of S.H.I.E.L.D. #33
  459. 459.0 459.1 Iron Man: Director of S.H.I.E.L.D. #34
  460. Nova Vol 4 #16
  461. 461.0 461.1 Nova #17
  462. 462.0 462.1 Guardians of the Galaxy Vol 2 #6
  463. Guardians of the Galaxy Vol 2 #4
  464. Guardians of the Galaxy Vol 2 #5
  465. Thunderbolts #124
  466. 466.0 466.1 Thunderbolts #125
  467. Secret Invasion: X-Men #1
  468. Secret Invasion: X-Men #2
  469. Secret Invasion: X-Men #3
  470. Secret Invasion: X-Men #4
  471. Secret Invasion: Thor #2
  472. Secret Invasion: Thor #3
  473. Nova Vol 4 #18
  474. She-Hulk Vol 2 #32
  475. New Warriors Vol 4 #15
  476. Deadpool Vol 4 #1
  477. Deadpool Vol 4 #2
  478. Deadpool Vol 4 #3
  479. 479.0 479.1 Secret Invasion #6
  480. Black Panther Vol 4 #39
  481. Black Panther Vol 4 #40
  482. Black Panther Vol 4 #41
  483. Iron Man: Director of S.H.I.E.L.D. #35
  484. Captain Britain and MI-13 #1
  485. Captain Britain and MI-13 #2
  486. Captain Britain and MI-13 #3
  487. Captain Britain and MI-13 #4
  488. Secret Invasion: Fantastic Four #3
  489. Incredible Hercules #119
  490. Secret Invasion #8
  491. 491.0 491.1 Skrull Kill Krew Vol 2 #5
  492. Dark Avengers #1
  493. New Avengers #48
  494. New Avengers #49
  495. Dark Reign: Elektra #1
  496. Dark Reign: Elektra #4
  497. Secret Invasion: War of Kings #1
  498. War of Kings: Ascension #1
  499. Secret Invasion Aftermath: Beta Ray Bill - The Green of Eden #1
  500. Guardians of the Galaxy Vol 2 #8
  501. Dark Reign: New Nation #1
  502. Skrull Kill Krew Vol 2 #2
  503. Skrull Kill Krew Vol 2 #3
  504. Skrull Kill Krew Vol 2 #4
  505. Spider-Woman Vol 4 #1
  506. Spider-Woman Vol 4 #3
  507. Spider-Woman Vol 4 #7
  508. Ms. Marvel Vol 2 #50
  509. Hawkeye & Mockingbird #6
  510. Annihilators #1
  511. New Avengers Annual Vol 2 #1
  512. Avengers Annual Vol 3 #1
  513. X-Men Vol 3 #28
  514. X-Men Vol 3 #29
  515. Avengers Vol 4 #30
  516. A + X #13
  517. A + X #14
  518. A + X #15
  519. A + X #16
  520. A + X #17
  521. A + X #18
  522. 522.0 522.1 522.2 Avengers Vol 5 #18
  523. 523.0 523.1 Avengers Vol 5 #15
  524. Infinity #1
  525. Infinity #2
  526. Avengers Vol 5 #19
  527. Infinity #3
  528. Avengers Vol 5 #20
  529. Infinity #4
  530. Avengers Vol 5 #21
  531. Infinity #5
  532. Avengers Vol 5 #22
  533. Avengers Vol 5 #23
  534. Infinity #6